河北唐山寒武系芙蓉统江山阶的Barbitositheca ansatus Qian et Jiang,1980
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摘要
本文描述了河北省唐山市丰润区寒武系芙蓉统江山阶长山组的锥石化石,鉴定为有柄琴状锥石(Barbitositheca ansatus Qian et Jiang, 1980)。化石保存为印模,轮廓及横肋特征清晰。其中一块标本保存有触手,是对该种特征的补充。目前的化石资料表明,寒武纪早期琴状锥石属主要分布在华南,寒武纪晚期分布范围扩大到华北,形态随时空发生一些变化,主要表现为华北标本角脊数量多于华南标本,并且前者每个角脊上都有一条触手。另外,研究发现寒武纪之后的锥石保存状态与寒武纪有所区别,可能反映了这类生物在不同时期内面临不同的生存压力。本文研究的寒武纪锥石为华北地区的首次发现,扩大了此类生物的时空分布。
This paper described Conulariids from the Changshan Formation, Jiangshanian Stage, Furongian Series(Cambrian) at Fengrun District in Tangshan city, Hebei Province. The fossils were identified as Barbitositheca ansatus Qian et Jiang, 1980. The fossils were preserved as impressions with clear outlines and transverse ribs. One of the specimens was preserved with tentacles, an important supplement to the characteristics of the species. The present fossil data indicates that Barbitositheca was mainly distributed in South China in the Early Cambrian, and then was extended to North China in the Late Cambrian. The specimen of Barbitositheca from North China holds three corner ridges, each of which owned one tentacle; While Barbitositheca from South China was reported to hold two corner ridges without tentacles. Observation on conulata also indicates that individuals from the Cambrian were preserved differently from those after Cambrian, which may be caused by different survival pressures. Herein it's the first time to report Cambrian conulata from North China, which expands the spatial and temporal distribution of this taxa.
This paper described Conulariids from the Changshan Formation, Jiangshanian Stage, Furongian Series(Cambrian) at Fengrun District in Tangshan city, Hebei Province. The fossils were identified as Barbitositheca ansatus Qian et Jiang, 1980. The fossils were preserved as impressions with clear outlines and transverse ribs. One of the specimens was preserved with tentacles, an important supplement to the characteristics of the species. The present fossil data indicates that Barbitositheca was mainly distributed in South China in the Early Cambrian, and then was extended to North China in the Late Cambrian. The specimen of Barbitositheca from North China holds three corner ridges, each of which owned one tentacle; While Barbitositheca from South China was reported to hold two corner ridges without tentacles. Observation on conulata also indicates that individuals from the Cambrian were preserved differently from those after Cambrian, which may be caused by different survival pressures. Herein it's the first time to report Cambrian conulata from North China, which expands the spatial and temporal distribution of this taxa.
引文
邢裕盛,丁启秀,罗惠麟等,1984.中国震旦系-寒武系界线,中国地质科学院地质研究所所刊,地质出版社.10:1-262
刘云焕,李勇,邵铁全等,2005.陕西南部早寒武世原始锥石类cari-nachitids两新种记述.微体古生物学报,22(3):311-321
刘云焕,李勇,邵铁全等,2011.陕西宁强原始锥石类五辐对称新属种的发现.微体古生物学报,28(2):244-249
朱志康,1985.中国泥盆纪和二叠纪锥石类新材料记叙.古生物学报,24(5):528-540
李声之,王继兴,王喜富等,1996.河北省岩石地层.武汉:中国地质大学出版社.1-146
杨暹和,何原相,邓守和,1983.四川南江地区震旦系-寒武系界线及小壳化石群.成都地矿所所刊,第4号:91-110
何原相,杨暹和,1986.四川南江早寒武世早期的腔肠动物化石.中国地质科学院成都地质矿产研究所所刊,第7号:31-48
何廷贵,1984.四川雷波牛牛寨下寒武统梅树村阶Lapworthella bella小壳化石组合的发现.地层古生物论文集,2:23-34
何廷贵,1987.扬子地台区早寒武世锥石动物及其早期演化.成都地质学院学报,14(2):7-20
陈孟莪,1982.四川峨眉麦迪平剖面震旦系-寒武系界线的新认识及有关化石群的记述.地质科学,3:253-264
张增奇,刘明渭,1996.山东省岩石地层.武汉:中国地质大学出版社.1-328
罗惠麟,蒋志文,武希彻等,1982.云南东部震旦系-寒武系界线.昆明:云南人民出版社.1-302
河北省地质矿产局,1989.河北省北京市天津市区域地质志.北京:地质出版社.1-741
徐桂荣,李凤麟,1979.锥石类新属种及其地层意义.地质学报,2:91-99
殷继成,丁莲芳,何廷贵等,1980.四川峨眉—甘洛地区震旦纪地层古生物及沉积环境.成都:四川人民出版社.1-231
钱逸,1977.华中西南区早寒武世梅树村阶的软舌螺纲及其它化石.古生物学报,16(2):255-227
钱逸,1999.中国小壳化石分类学与生物地层学.北京:科学出版社.1-247
彭善池,2009.华南斜坡相寒武纪三叶虫动物群研究回顾并论我国南、北方寒武系的对比.古生物学报,48(3):437-452
蒋志文,1980.云南晋宁梅树村及梅树村动物群.中国地质科学院院报,2(1):75-92
BUREAU of Geology and Mineral Resources of Hebei Province,1989.Regional Geology of Hebei Province,Beijing City,and Tianjin City.Beijing:Geological Publishing House.1-741(in Chinese)
BABCOCK L E,FELDMANN R M,WILSON M T,1987.Teratology and pathology of some Paleozoic conulariids.Lethaia,20:93-105
BRASIER M D,SINGH P,1987.Microfossils and precambrian cambrian boundary stratigraphy at Maldeota(spacing),Lesser Himalaya.Geol.Mag.,124(4):323-345
BROOD K,1995.Morphology,structure,and systematics of the conulariids.GFF,117(3):121-137
CHEN Menge,1982.The new knowledge of the fossil assembages from Maidiping Section,Emei county,Sichuan with reference to the Sinian-Cambrian Boundary.Scientia Geologica Sinica,3:253-264
DUAN Baichuan,DONG Xiping,PORRAS L et al.,2017.The early Cambrian fossil embryo Pseudooides is a direct-developing cnidarian,not an early ecdysozoan.The Royal Society Publishing.1-10
FATKA O,KRAFT P,SZABAD M,2012.A first report of Sphenothallus Hall,1847 in the Cambrian of Variscan Europe.Comptes Rendus Palevol,11:539-547
HE Yuanxiang,YANG Xianhe,1986.Early Cambrian coelenterates from Nanjiang,Sichuang.Bull.Chengdu Inst.Geol.M.R.,Chinese Acad.Geol.Sci.,7:31-48(in Chinese with English abstract)
HE Tinggui,1984.Discovery of Lapworthella bella assemblage from Lower Cambrain Meishucun Stage in Niuniuzhai,Leibo county,Sichuan Province.Collection of Papers on Stratigraphic Paleontology,2:23-34(in Chinese with English abstract)
HE Tinggui,1987.Early Cambrian conulariids from Yangtze Platform and their early evolution.Journal of Chengdu College of Geology,14(2):7-20(in Chinese with English abstract)
HUGHES N C,GUNDERSON G O,WEEDON M J,2000.Late Cambrian conulariids from Wisconsin and Minnesota.Journal of Paleontology,74:828-839
IVANTSOV A Y,FEDONKIN M A,2002.Conulariid-like fossil from the Vendian of Russia;a metazoan clade across the Prote-rozoic/Palaeozoic boundary.Palaeontology,45:1219-1229
JIANG Zhiwen,1980.The Meishucun Stage and Fauna of the Jinning County,Yunnan.Bulletin Chinese Acad.Geol.Sci.Series I,2(1):75-92(in Chinese with English abstract)
KIDERLEN I,1937.Die Conularien.Uber Bau und leben der ersten Scyphozoa,Neues Jahrbuch Für Mineralogie,usw,Beil,77,Abt.B:113-169
LUO Huilin,JIANG Zhiwen,WU Xiche et al.,1982.The Sinan-Cambrian boundary in Eastern Yunnan,China.Kunming:Yunnan Institute of Geological Sciences.1-302(in Chinese)
LUO Huilin,JIANG Zhiwen,WU Xiche et al.,1984.Sinian-Cambrian Boundary Stratotype Section at Meishucun,Jinning,Yunnan,China.Kunming:Yunnan People's Publishing House.154(in Chinese)
LI Shengzhi,WANG Jixing,WANG Xifu et al.,1996.Stratigraphy(Lithostratic) of Hebei Province.Wuhan:China University of Geosciences Press.1-146(in Chinese)
LIU Yunhuan,LI Yong,SHAO Tiequan et al.,2005.Two new species of protoconulariids from the Early Cambrian in south Shaanxi,China.Acta Micropalaeontologica Sinica,22(3):311-321(in Chinese with English abstract)
LIU Yunhuan,LI Yong,SHAO Tiequan et al.,2011.A new genus and species of protoconulariids from early Cambrian in Southern Shaanxi,China.Acta Micropalaeontologica Sinica,28(2):244-249(in Chinese with English abstract)
LEME J D M,SIM?ES M G,MARQUES A C,2008.Cladistic analysis of the suborder conulariina miller and gurley,1896(Cnidaria,Scyphozoa;Vendian-Triassic).Palaeontology,51(3):649-662
LUCAS S G,2012.The extinction of the conulariids.Geosciences,2:1-10
PENG Shanchi,2009.Review on the studies of Cambrian trilobite faunas from Jiangnan slope belt,South China,with notes on Cambrian correlation between South and North China.Acta Palaeontologica Sinica,48(3):437-452(in Chinese with English abstract)
QIAN Yi,1977.Hyolitha and some problematica from the Lower Cambrian Meishucun Stage in Central and SW China,Acta Palaeontologica Sinica,16(2):255-275(in Chinese with English summary)
QIAN Yi,1999.Taxonomy and Biostratigraphy of Small Shelly Fossils in China.Beijing:Science Press.22-117
SENDINO C,ZáGOR?EK K,VYHLASOVá Z,2011.The aperture and its closure in an Ordovician conulariid.Acta Palaeontologica Polonica,56(3):659-663
VAN ITEN H,COX R S,1992.Evidence of clonal budding in a radial cluster of Paraconularia crustula (White) (Pennsylvanian: Cnidaria).Lethaia,25:421-426
VAN ITEN H,LEME J D M,RODRIGUES S C et al.,2005.Reinterpretation of a conulariid-like fossil from the Vendian of Russia.Palaeontology,48:619-622
VAN ITEN H,LEME J D M,RODRIGUES S C et al.,2006.New data on the anatomy of Conularia milwaukeensis Cleland,1911(Middle Devonian,Iowa and Wisconsin).Journal of Paleonto-logy,80(2):392-395
VAN ITEN H,SüDKAMP W H,2010.Exceptionally preserved conulariids and an edrioasteroid from the Hunsrück slate(lower Devonian,SW Germany).Palaeontology,53(2):403-414
VAN ITEN H,VICTOR J R,STRAETEN C A V et al.,2013.Life mode of in situ conularia in a middle Devonian Epibole.Palaeontology,56(1):29-48
VAN ITEN H,MUIR L,SIM?E M G et al.,2016.Palaeobio-geography,palaeoecology and evolution of Lower Ordovician conulariids and Sphenothallus (Medusozoa,Cnidaria),with emphasis on the Fezouata Shale of southeastern Morocco.Palaeo-geography,Palaeoclimatology,Palaeoecology,460:170-178
WATERHOUSE J B,1979.Permian and Triassic conulariid species from New Zealand.Journal of the Royal Society of New Zea-land,9:475-489
XING Yusheng,DING Qixiu,LUO Huilin et al.,1984.The Sinian-Cambrian Boundary of China.Bulletin of the Institute of Geology,Chinese Academy of Geological Science (Special Issue on the Sinian-Cambrian Boundary of China),10:1-262(in Chinese with English summary)
XU Guirong,LI Fenglin,1979.New conulariids genera and species in China.Acta Geologica Sinica,2:91-99(in Chinese with English summary)
YIN Jicheng,DING Lianfang,HE Tinggui et al.,1980.The Palaeontology and Sedimentary Environment of the Sinian System in Emei-Ganluo Area.Sichuan:Sichuan People's Publishing Hou-se.1-231(in Chinese with English summary)
YANG Xianhe,HE Yuanxiang,DENG Shouhe,1983.On the Sinian Cambrian Boundary and the Small Shelly Fossil Assemblages in Nanjiang Area,Sichuan.Bulletin of the Chengdu Institute of Geology and Mineral Resources,Chinese Academy of Geological Science,4:91-110(in Chinese with English summary)
ZHANG Zengqi,LIU Mingwei,1996.Rock formations in Shandong Province.Wuhan:China University of Geosciences Press.1-328(in Chinese)
ZHU Zhikang,1985.New materials of Devonian and Permian conulariids from China.Acta Palaeontologica Sinica,24(5):528-540(in Chinese with English abstract)
刘云焕,李勇,邵铁全等,2005.陕西南部早寒武世原始锥石类cari-nachitids两新种记述.微体古生物学报,22(3):311-321
刘云焕,李勇,邵铁全等,2011.陕西宁强原始锥石类五辐对称新属种的发现.微体古生物学报,28(2):244-249
朱志康,1985.中国泥盆纪和二叠纪锥石类新材料记叙.古生物学报,24(5):528-540
李声之,王继兴,王喜富等,1996.河北省岩石地层.武汉:中国地质大学出版社.1-146
杨暹和,何原相,邓守和,1983.四川南江地区震旦系-寒武系界线及小壳化石群.成都地矿所所刊,第4号:91-110
何原相,杨暹和,1986.四川南江早寒武世早期的腔肠动物化石.中国地质科学院成都地质矿产研究所所刊,第7号:31-48
何廷贵,1984.四川雷波牛牛寨下寒武统梅树村阶Lapworthella bella小壳化石组合的发现.地层古生物论文集,2:23-34
何廷贵,1987.扬子地台区早寒武世锥石动物及其早期演化.成都地质学院学报,14(2):7-20
陈孟莪,1982.四川峨眉麦迪平剖面震旦系-寒武系界线的新认识及有关化石群的记述.地质科学,3:253-264
张增奇,刘明渭,1996.山东省岩石地层.武汉:中国地质大学出版社.1-328
罗惠麟,蒋志文,武希彻等,1982.云南东部震旦系-寒武系界线.昆明:云南人民出版社.1-302
河北省地质矿产局,1989.河北省北京市天津市区域地质志.北京:地质出版社.1-741
徐桂荣,李凤麟,1979.锥石类新属种及其地层意义.地质学报,2:91-99
殷继成,丁莲芳,何廷贵等,1980.四川峨眉—甘洛地区震旦纪地层古生物及沉积环境.成都:四川人民出版社.1-231
钱逸,1977.华中西南区早寒武世梅树村阶的软舌螺纲及其它化石.古生物学报,16(2):255-227
钱逸,1999.中国小壳化石分类学与生物地层学.北京:科学出版社.1-247
彭善池,2009.华南斜坡相寒武纪三叶虫动物群研究回顾并论我国南、北方寒武系的对比.古生物学报,48(3):437-452
蒋志文,1980.云南晋宁梅树村及梅树村动物群.中国地质科学院院报,2(1):75-92
BUREAU of Geology and Mineral Resources of Hebei Province,1989.Regional Geology of Hebei Province,Beijing City,and Tianjin City.Beijing:Geological Publishing House.1-741(in Chinese)
BABCOCK L E,FELDMANN R M,WILSON M T,1987.Teratology and pathology of some Paleozoic conulariids.Lethaia,20:93-105
BRASIER M D,SINGH P,1987.Microfossils and precambrian cambrian boundary stratigraphy at Maldeota(spacing),Lesser Himalaya.Geol.Mag.,124(4):323-345
BROOD K,1995.Morphology,structure,and systematics of the conulariids.GFF,117(3):121-137
CHEN Menge,1982.The new knowledge of the fossil assembages from Maidiping Section,Emei county,Sichuan with reference to the Sinian-Cambrian Boundary.Scientia Geologica Sinica,3:253-264
DUAN Baichuan,DONG Xiping,PORRAS L et al.,2017.The early Cambrian fossil embryo Pseudooides is a direct-developing cnidarian,not an early ecdysozoan.The Royal Society Publishing.1-10
FATKA O,KRAFT P,SZABAD M,2012.A first report of Sphenothallus Hall,1847 in the Cambrian of Variscan Europe.Comptes Rendus Palevol,11:539-547
HE Yuanxiang,YANG Xianhe,1986.Early Cambrian coelenterates from Nanjiang,Sichuang.Bull.Chengdu Inst.Geol.M.R.,Chinese Acad.Geol.Sci.,7:31-48(in Chinese with English abstract)
HE Tinggui,1984.Discovery of Lapworthella bella assemblage from Lower Cambrain Meishucun Stage in Niuniuzhai,Leibo county,Sichuan Province.Collection of Papers on Stratigraphic Paleontology,2:23-34(in Chinese with English abstract)
HE Tinggui,1987.Early Cambrian conulariids from Yangtze Platform and their early evolution.Journal of Chengdu College of Geology,14(2):7-20(in Chinese with English abstract)
HUGHES N C,GUNDERSON G O,WEEDON M J,2000.Late Cambrian conulariids from Wisconsin and Minnesota.Journal of Paleontology,74:828-839
IVANTSOV A Y,FEDONKIN M A,2002.Conulariid-like fossil from the Vendian of Russia;a metazoan clade across the Prote-rozoic/Palaeozoic boundary.Palaeontology,45:1219-1229
JIANG Zhiwen,1980.The Meishucun Stage and Fauna of the Jinning County,Yunnan.Bulletin Chinese Acad.Geol.Sci.Series I,2(1):75-92(in Chinese with English abstract)
KIDERLEN I,1937.Die Conularien.Uber Bau und leben der ersten Scyphozoa,Neues Jahrbuch Für Mineralogie,usw,Beil,77,Abt.B:113-169
LUO Huilin,JIANG Zhiwen,WU Xiche et al.,1982.The Sinan-Cambrian boundary in Eastern Yunnan,China.Kunming:Yunnan Institute of Geological Sciences.1-302(in Chinese)
LUO Huilin,JIANG Zhiwen,WU Xiche et al.,1984.Sinian-Cambrian Boundary Stratotype Section at Meishucun,Jinning,Yunnan,China.Kunming:Yunnan People's Publishing House.154(in Chinese)
LI Shengzhi,WANG Jixing,WANG Xifu et al.,1996.Stratigraphy(Lithostratic) of Hebei Province.Wuhan:China University of Geosciences Press.1-146(in Chinese)
LIU Yunhuan,LI Yong,SHAO Tiequan et al.,2005.Two new species of protoconulariids from the Early Cambrian in south Shaanxi,China.Acta Micropalaeontologica Sinica,22(3):311-321(in Chinese with English abstract)
LIU Yunhuan,LI Yong,SHAO Tiequan et al.,2011.A new genus and species of protoconulariids from early Cambrian in Southern Shaanxi,China.Acta Micropalaeontologica Sinica,28(2):244-249(in Chinese with English abstract)
LEME J D M,SIM?ES M G,MARQUES A C,2008.Cladistic analysis of the suborder conulariina miller and gurley,1896(Cnidaria,Scyphozoa;Vendian-Triassic).Palaeontology,51(3):649-662
LUCAS S G,2012.The extinction of the conulariids.Geosciences,2:1-10
PENG Shanchi,2009.Review on the studies of Cambrian trilobite faunas from Jiangnan slope belt,South China,with notes on Cambrian correlation between South and North China.Acta Palaeontologica Sinica,48(3):437-452(in Chinese with English abstract)
QIAN Yi,1977.Hyolitha and some problematica from the Lower Cambrian Meishucun Stage in Central and SW China,Acta Palaeontologica Sinica,16(2):255-275(in Chinese with English summary)
QIAN Yi,1999.Taxonomy and Biostratigraphy of Small Shelly Fossils in China.Beijing:Science Press.22-117
SENDINO C,ZáGOR?EK K,VYHLASOVá Z,2011.The aperture and its closure in an Ordovician conulariid.Acta Palaeontologica Polonica,56(3):659-663
VAN ITEN H,COX R S,1992.Evidence of clonal budding in a radial cluster of Paraconularia crustula (White) (Pennsylvanian: Cnidaria).Lethaia,25:421-426
VAN ITEN H,LEME J D M,RODRIGUES S C et al.,2005.Reinterpretation of a conulariid-like fossil from the Vendian of Russia.Palaeontology,48:619-622
VAN ITEN H,LEME J D M,RODRIGUES S C et al.,2006.New data on the anatomy of Conularia milwaukeensis Cleland,1911(Middle Devonian,Iowa and Wisconsin).Journal of Paleonto-logy,80(2):392-395
VAN ITEN H,SüDKAMP W H,2010.Exceptionally preserved conulariids and an edrioasteroid from the Hunsrück slate(lower Devonian,SW Germany).Palaeontology,53(2):403-414
VAN ITEN H,VICTOR J R,STRAETEN C A V et al.,2013.Life mode of in situ conularia in a middle Devonian Epibole.Palaeontology,56(1):29-48
VAN ITEN H,MUIR L,SIM?E M G et al.,2016.Palaeobio-geography,palaeoecology and evolution of Lower Ordovician conulariids and Sphenothallus (Medusozoa,Cnidaria),with emphasis on the Fezouata Shale of southeastern Morocco.Palaeo-geography,Palaeoclimatology,Palaeoecology,460:170-178
WATERHOUSE J B,1979.Permian and Triassic conulariid species from New Zealand.Journal of the Royal Society of New Zea-land,9:475-489
XING Yusheng,DING Qixiu,LUO Huilin et al.,1984.The Sinian-Cambrian Boundary of China.Bulletin of the Institute of Geology,Chinese Academy of Geological Science (Special Issue on the Sinian-Cambrian Boundary of China),10:1-262(in Chinese with English summary)
XU Guirong,LI Fenglin,1979.New conulariids genera and species in China.Acta Geologica Sinica,2:91-99(in Chinese with English summary)
YIN Jicheng,DING Lianfang,HE Tinggui et al.,1980.The Palaeontology and Sedimentary Environment of the Sinian System in Emei-Ganluo Area.Sichuan:Sichuan People's Publishing Hou-se.1-231(in Chinese with English summary)
YANG Xianhe,HE Yuanxiang,DENG Shouhe,1983.On the Sinian Cambrian Boundary and the Small Shelly Fossil Assemblages in Nanjiang Area,Sichuan.Bulletin of the Chengdu Institute of Geology and Mineral Resources,Chinese Academy of Geological Science,4:91-110(in Chinese with English summary)
ZHANG Zengqi,LIU Mingwei,1996.Rock formations in Shandong Province.Wuhan:China University of Geosciences Press.1-328(in Chinese)
ZHU Zhikang,1985.New materials of Devonian and Permian conulariids from China.Acta Palaeontologica Sinica,24(5):528-540(in Chinese with English abstract)
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