中国树甲族分类研究（鞘翅目：拟步甲科）
|
摘要
树甲族Strongyliini昆虫是较为重要的森林害虫,隶属于鞘翅目Coleoptera、多食亚目Polyphaga、拟步甲总科Tenebrionoidea、拟步甲科Tenebrionidae、圆甲亚科Coelometopinae。目前全球已记述60属约2200种(Masumoto,1996)。本研究首次对我国树甲族种类进行较全面的总结,内容包括总论和各论两部分。
总论部分详细介绍了树甲族的研究概况、分类地位与分类系统、材料与方法、形态特征以及生物学特性和经济意义。并首次对2属31种树甲后翅进行了解剖和摄像。通过比较研究发现,翅脉在属内差异小,属间差异较大,主要表现在1A脉和2A脉是否具有共同的分节点。树甲属中存在这一节点,而在铜树甲中两脉通过1条短横脉相连,并且2A脉与3A脉也有1短脉相连。另外对我国树甲种类的地理分布进行了初步分析。
各论部分记述中国树甲族昆虫2属84种(亚种),其中包括9新种,1中国新记录种,2雄性新发现和1疑问种。新种为:云南铜树甲Ainu yunnanense sp. nov.,扶隆铜树甲A. fulongium sp. nov.,道真树甲Strongylium daozhenense sp. nov.,武夷树甲S. wuyishanense sp. nov.,直翅树甲S. orthopterus sp. nov.,短胸树甲S. brachythorax sp. nov.,尖须树甲S. oxypalus sp. nov.,红翅树甲S. erythroelytrae sp. nov.,短头树甲S. brachycephalic sp. nov.;新记录种为:波形树甲S. undulatum Fairmaire,1903。
The tribe Strongyliini, belonging to the family Tenebrionidae of the order Coleoptera , is of rather great economic importance, and comprises more than 2200 species belonging to 60 genera all over the world (Masumoto, 1996). The species of the tribe Strongyliini from China are firstly delt with in this present paper, which comprises the general account and the systematic account.In the general account a classificatory review, the status of the classification and hierarchy of the tribe Strongyliini are made, and the morphological characteristics, biology and economic importance are also introduced in detail. The hind wings of 31 species belonging to 2 genera are scissored and photographed, the research shows that the difference of the nervure between 2 genera is more obvious than among species in a genus. Besides, the species' distribution of the tribe Strongyliini in China is also presented and discussed.In the systematic account 84 species known to China belonging to 2 genera are described in detail. Of them, 9 species are new to science: Ainu yunnanense sp. nov., A. fulongium sp. nov., Strongylium daozhenense sp. nov., S. wuyishanense sp. nov., S. orthopterus sp. nov., S. brachythorax sp. nov., S. oxypalus sp. nov., S. erythroelytrae sp. nov., S. brachycephalic sp. nov.; a species is newly recorded from China: S. undulatum Fairmaire, 1903, in addition, the male of 2 species are firstly recorded.
总论部分详细介绍了树甲族的研究概况、分类地位与分类系统、材料与方法、形态特征以及生物学特性和经济意义。并首次对2属31种树甲后翅进行了解剖和摄像。通过比较研究发现,翅脉在属内差异小,属间差异较大,主要表现在1A脉和2A脉是否具有共同的分节点。树甲属中存在这一节点,而在铜树甲中两脉通过1条短横脉相连,并且2A脉与3A脉也有1短脉相连。另外对我国树甲种类的地理分布进行了初步分析。
各论部分记述中国树甲族昆虫2属84种(亚种),其中包括9新种,1中国新记录种,2雄性新发现和1疑问种。新种为:云南铜树甲Ainu yunnanense sp. nov.,扶隆铜树甲A. fulongium sp. nov.,道真树甲Strongylium daozhenense sp. nov.,武夷树甲S. wuyishanense sp. nov.,直翅树甲S. orthopterus sp. nov.,短胸树甲S. brachythorax sp. nov.,尖须树甲S. oxypalus sp. nov.,红翅树甲S. erythroelytrae sp. nov.,短头树甲S. brachycephalic sp. nov.;新记录种为:波形树甲S. undulatum Fairmaire,1903。
The tribe Strongyliini, belonging to the family Tenebrionidae of the order Coleoptera , is of rather great economic importance, and comprises more than 2200 species belonging to 60 genera all over the world (Masumoto, 1996). The species of the tribe Strongyliini from China are firstly delt with in this present paper, which comprises the general account and the systematic account.In the general account a classificatory review, the status of the classification and hierarchy of the tribe Strongyliini are made, and the morphological characteristics, biology and economic importance are also introduced in detail. The hind wings of 31 species belonging to 2 genera are scissored and photographed, the research shows that the difference of the nervure between 2 genera is more obvious than among species in a genus. Besides, the species' distribution of the tribe Strongyliini in China is also presented and discussed.In the systematic account 84 species known to China belonging to 2 genera are described in detail. Of them, 9 species are new to science: Ainu yunnanense sp. nov., A. fulongium sp. nov., Strongylium daozhenense sp. nov., S. wuyishanense sp. nov., S. orthopterus sp. nov., S. brachythorax sp. nov., S. oxypalus sp. nov., S. erythroelytrae sp. nov., S. brachycephalic sp. nov.; a species is newly recorded from China: S. undulatum Fairmaire, 1903, in addition, the male of 2 species are firstly recorded.
引文
黑佐,福田,林长闲.1976.日本幼虫图鉴.东京:图鉴北隆馆.
李鸿兴.1997.鞘翅目.拟步甲科.杨星科主编.1997.长江三峡库区昆虫.重庆:重庆出版社,693.
刘晓丽.2004.部分拟步甲线粒体16S rDNA和18S rDNA基因序列与分子系统学研究[研究生毕业论文].河北大学.
任国栋,杨秀娟,印红.2002.基于防御腺的中国琵甲族Blaptini系统发育分析(鞘翅目:拟步甲科).昆虫学报,45(6):805-811.
任国栋,于有志.1999.中国荒漠半荒漠地区拟步甲分类研究.保定:河北大学出版社.
任国栋.1998.鞘翅目.拟步甲科.吴鸿主编.1998.龙王山昆虫.北京:中国林业出版社,111.
任国栋.2002.拟步甲科.黄邦侃主编.2002.福建昆虫志.第六卷.福州:福建科学技术出版社,137-170.
杨星科.1999.鞘翅目.郑乐怡,归鸿主编.1999.昆虫分类.南京:南京师范大学出版社,573.
叶晔,蒋鹏,任国栋.2003.31属拟步甲的防御腺特征与系统发育关系初析.河北大学学报(自然科学版),23(3):279-292.
于洋.2005.拟步甲成虫口器比较形态学研究(一).河北大学学报(自然科学版),25(3).(发表中).
张荣祖.1998.“中国动物地理区划”的再修订.动物分类学报,23(增刊):207-222.
张荣祖.1999.中国动物地理.北京:科学出版社.
赵养昌.1963.中国经济昆虫志.第四册.鞘翅目.伪步行虫科.北京:科学出版社.
周红章.2002.关于亚种下名称的可用性问题:实例与分析.动物分类学报,27(4):659-664.
周红章.2003.种级异物同名处理及属名变动影响分析.动物分类学报,28(1):5-12.
Ando, K. 2003. A new Strongylium species (Coleoptera:Tenebrionidae) occurring in Japan. Ent. Rev. Japan, 58(1):79-81, figs.1-2.
Arnett, R. H. Jr. 1968. The beetles of the United States (A manual for identification). Catholic University of America Press, Washington, D. C., 1112.
Bouchard, P. 2002. Phylogenetic revision of the flightless Australian genus Apterotheca Gebien (Coleoptera:Tenebrionidae: Coelometopinae). Invertebrate systematics,16: 449-554.
Bouchard, P., W. E. Steiner Jr. 2004. First descriptions of Coelometopini pupae (Coleoptera: Tenebrionidae) from Australia, Southeast Asia and the Pacific region, with comments on phylogenetie relationships and antipredator adaptations. Systematic Entomology, 29:101-102.
Chujo, M. 1968a. A check list of the Formosa Tenebrionidae (Coleoptera) Ⅱ. Kontyu, 36(1): 15-17, fig. 1.
Chujo, M. 1968b. A check list of the Formosa Tenebrionidae (Coleoptera) Ⅲ. Kontyu, 36(4): 397.
Chujo, M. 1978. Tenebrionidae of the Nansei Islands Ⅲ (Coleoptera). Esakia, (11): 78-80.
Chujo, M. 1979. Tenebrionidae of the Nansei Islands Ⅴ (Coleoptera). Esakia, (14): 54-56.
Chujo, M. 1998. Tenebrionidae (except genera Platydema Ischnodactylus, tribe Cnodalonini). In: Kurosawa, Y., Hisamatsu, S., Sasaji, H. 1998. the Coleoptera of Japan in color Vol. Ⅲ. Japan: Hoikusha Publishing Co., Ltd., 338-339.
Chujo, M., C. E. Lee. 1992. Tenebrionidae from Chejudo Island, Korea (Insecta, Coleoptera). Esakia, 32: 43-44, fig. 11.
Chujo, M., C. E. Lee. 1993. Korean Tenebrionidae (Insecta, Coleoptera). Esakia, 33:120. fig. 12,
Doyen, J. T. 1972. Familial and subfamilial classification of the Tenebrionidea (Coleoptera) and a revised generic classification of the Coniontini (Tentyriidae).Quaesitiones. Ent., 8(4): 357-367.
Doyen J. T. 1982. Phenetic and cladistie relationships among Tenebrionid beetles (Coleoptera). Syst. Entomol., 7(2): 127-183.
Doyen J. T. 1989. Reeonstitutions of Coelometopini Tenebrionini and related tribes of America north of Colombia (Coleoptera:Tenebrionidae). Journal of the New York Entomological Society, 97(3): 300-301, figs. 25-26.
Doyen J. T., J. F. Lawrence. 1979. Relationships and higher classification of some Tenebrionidae and Zopheridae (Coleoptera). Syst. Entomol., 4(4): 333-377.
Doyen J. T., R. E. Somerby. 1974. Phenetic similarity and mullerian mimicry among darking ground beetles(Coleoptera:Tenebrionidae).The Canadian Entomologist, 106: 771-772.
Fairmaire, L. 1891. Comptes-rendus des seances. Annales de la Societe Entomologique de Belgique, T., 35: 212-214.
Fairmaire, L. 1893. Coleopteres du Haut Tonkin. Annales de la Societe Entomologique de Belgique.,37: 319-321.
Fairmaire, L. 1893. Heteromeres. Annales de la Societe Entomologique de Belgique, 37:319-320.
Fairmaire, L. 1896. Note XII Coleopteres de L'inde Boreale, China et Malaisie. Notes from the Leyden Museum, 18:112-115.
Fairmaire, L. 1902. Descriotions de Coleopteres recueillis en Chine par M. de Latouche. Bulletin de la Societe Entomologique de France, 318.
Fairmaire, L. 1903. Descriptions de Quelques Heteromeres. Annales de la Societe Entomologique de Belgique, 47: 18-20.
Fairmaire, L., 1893. Note sur quelques Coleopteres des environs de Lang-Song. Annales de la Societe Entomologique de Belgique, 37: 299.
Gebien, H. 1913a. Die Tenebrionidae der Philippinen. The Philippine Journal of Science, 8(6):431-432.
Gebien, H. 1913b. H. Sauter's Formosa-A usbeute. Tenebrionidae (Coleoptera), Archiv fur Naturgeschichte, 79(A9): 44-54, figs. 1, 3, 4.
Gebien, H. 1944. Katalog der Tenebrioniden (Coleoptera) Ⅳ. Mitteilungen der Munchener Entomologischen Gesellschaft, 34:514(859)-555(899).
Imasaka, S. 1985. On the distributions of Strongylium marseuli Lewis and Strongylium cultellatum Maklin. Gekkan-Mushi, 172: 23-25.
Kaszab, Z. 1941. Tenebrioniden aus Formosa (Coleoptera). Stettiner Entomologische Zeitung, 102:71-72.
Kaszab, Z. 1954. Tenebrioniden Collected by J. Klapperich in Fukien. Ann. hist.-nat. Mus. Hung. (N. S.), 5: 247-264, figs.
Kaszab, Z. 1957. Neue Tenebrioniden (Coleoptera) aus Fukien. Bonn. Zool. Beitr., Bonn, 8(1): 56-63.
Kaszab, Z. 1964. Uber die Tenebrioniden einiger Japanischen Inseln (Ⅱ) (Coleoptera). Ent. Rev. Japan, Osaka,17: 8-10.
Kaszab, Z. 1977. Die Tenebrioniden des Papuanischen Gebietes Ⅰ. Strongyliini (Coleoptera: Tenebrionidae). Pacific Insects Monograph, 33: 1-219.
Kaszab, Z. 1980. Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici. 72:181.
Kulzer, H. 1966. Australische und Papuanische Strongyliini 29. Beitrag zur Kenntnis der Tenebrioniden. Ent. Arb. Mus. Frey, 338-353.
Kwon, Y. J., Y. S. Choi. 1986: Check list of Family Tenebrionidae from Korea. Insecta Koreana, 6:112-113
Lewis, G. 1894. On the Tenebrionidae of Japan. Annals & Magazine Natural History, 6(13): 479-483.
Maklin, F. W. 1864. Monographie der Gattung Strongylium Kirby, Lacordaire und der damit zunachst verwandten Formen. 518 pp., 4 pls. Finnlandischen Wissenschaftlichen Gesellschsft, Helsingfors.
Masumoto, K. 1981a. Tenebrionidae of Formosa (1). Elytra, 8(2): 37-53, figs. 1-20.
Masumoto, K. 1981b. Tenebrionidae of Formosa (2). Elytra, 9(1): 15-18, figs. 1-2.
Masumoto, K. 1981c. Tenebrionidae of Formosa (3). Elytra, 9(2): 95, 97, figs. 13-14.
Masumoto, K. 1981d. Notes and descriptions of Japanese Tenebrionidae (Ⅰ). Ent. Rev. Japan, 35:29-31, figs. 1-2.
Masumoto, K. 1982. Tenebrionidae of Formosa (5). Elytra, 10(2): 53-56, figs. 1, 1a, b, c, 2, 2a, b, c.
Masumoto, K. 1992. Tenebrionidae of East Asia (Ⅷ), Three new species of the genera Campsiomorpha (Cnodalonini) and Strongylium (Strongyliini) from Taiwan. Elytra, 20(1): 92-94, figs. 3, 4, 9-10.
Masumoto,K.1995.比和町立自然科学博物馆所藏台湾产科甲虫.(安藤+).比和科学 博物馆研究报告,33:17-22.
Masumoto, K. 1996a. New Tenebrionid beetles of the tribes Strongyliini, Misolampini and Adeliini (Coleoptera) from Northern Vietnam. Bull. Natn. Sci. Mus., Tokyo, (A), 22: 33-43.
Masumoto, K. 1996b. Replacement of the preoccupied name of a Strongylium species, Ent. Rev. Japan, 51: 14.
Masumoto, K. 1996c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅰ. Six new Strongylium species from Thailand, Laos and Taiwan, together with a new replacement name. Elytra, Tokyo, 24(1): 131-140, figs. 1-8.
Masumoto, K. 1996d. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅱ. New Strongylium species from Northern Thailand (Part 1). Elytra, Tokyo, 24(2): 337-366, figs. 1-20.
Masumoto, K. 1997a. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅲ. New Strongylium species from Southern India, preserved in the collections of the Museum National d'Histoire Naturelle, Paris and the Natural History Museum, London. Elytra, Tokyo, 25(1): 45-65, figs. 1-16.
Masumoto, K. 1997b. New records of Strongylium (Coleoptera,Tenebrionidae) from Northern Thailand. Elytra, Tokyo, 25(2): 65-66.
Masumoto, K. 1997c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅳ. Three species-groups of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 25(2): 279-309, figs. 1-56.
Masumoto, K. 1997d. New synonymy in the genus Strongylium (Coleoptera, Tenebrionidae, Strongyliini) described by Fairmaire and Pic. Elytra, Tokyo, 25(2): 309-310.
Masumoto, K. 1998a. New synonymy in the genus Strongylium (Coleoptera, Tenebrionidae, Strongyliini). Elytra, Tokyo, 26(1):200.
Masumoto, K. 1998b. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) V. Twenty new species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 26(1): 173-200, figs. 1-56.
Masumoto, K. 1998c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅵ. New Strongylium species from Northern Thailand (Part 2). Elytra, Tokyo, 26(2): 403-432, figs. 1-49.
Masumoto, K. 1999a. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅶ. Brachypterous strongyliines. Elytra, Tokyo, 27(1): 113-125, figs. 1-23.
Masumoto, K. 1999b. The male of Strongylium nanfangum (Coleoptera,Tenebrionidae). Elytra, Tokyo, 27(1):127-129.
Masumoto, K. 1999c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae)Ⅷ.Ten new species of the genus Stongylium.Elytra,Tokyo, 27(2): 335-352, figs.1-31.
Masumoto, K. 2000a. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) IX. Hairy Strongylium species from Southeast Asia (part Ⅰ. Species-group of Strongylium gibbosulum). Elytra, Tokyo, 28(1): 163-172, figs. 1-17.
Masumoto, K. 2000b. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) X. Ten new species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 28(2): 391-407, figs. 1-31.
Masumoto, K. 2001. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) XI. Ten new small-sized species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 29(2): 401-418, figs. 1-30.
Masumoto, K. 2002. A new Strongylium (Coleoptera,Tenebrionidae, Strongyliini) from Borneo. Elytra, Tokyo, 30(2): 335-338. figs. 1-3.
Masumoto, K. 2003. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅻ. New Strongylium species from Thailand. Elytra, 31(1): 65-86.
Medvedev, G. S. 1977. Taxonomical study of antennal sensillia in Tenebrionidae (Coleoptera: Tenebrionidae). Trudy Vses. Ent. Obshch, 58: 61-86, illust. [in Russian]
Miwa, Y. 1931. A systematic catalogue of Formosan Coleoptera. Rept. 55 Dept. Agr. Gov. Res. Inst. Formosa, 166.
Miwa, Y. 1939. Descriptions on four new species belonging to the family Tenebrionidae from Formosa. Zoological Magazine, Tokyo, 51(7): 414-415, figs. 3-4.
Nakane, T. 1968. New or little-known Coleoptera from Japan and its adjacent regions 27. Fragmenta Coleopterol., 21: 83-85.
Nomura, S. 1963: Some new species of the Coleoptera from Loochoo Is. and its adjacent regions, The Entpmological Review of Japan, 15(2): 37-40; P1.5, figs. 1-4.
Nomura, S. 1964: Some new species of the Coleoptera from Loochoo Is. and its Adjacent Regions, Ⅱ. The Entpmological Review of Japan, 17(2): 47-49. P1.3, fig. 4.
Pic, M. 1916. Especes et varietes nouvelles. Melanges Exotico-Entomologiques, 19: 12-13.
Pic, M. 1917. Coleopteres exotiques en partie nouveaux. L'Echange, Revue Linneenne, 33: 12.
Pic, M. 1928. Cinquantr-et-unieme fascicule. Melanges Exotico Entomologiques, 51: 23-24.
Pic, M. 1936. Nouveautes diverses. Melanges Exotico Entomologiques, 68: 19-20.
Pic, M. 1940. Diagnoses de Coleopteres exotiques. L'Echange, Revue Linneenne, 56:13.
Reitter, E. 1904. Bestimmungs-tabelle Tenebrioniden-unterfanilien: Lachnogyini, Akidini, Opatrini und Trachyscelini aus Europa und den angrenzenden Landern. Verh. Nat.-forsch. Ver. Brunn,12:25-189.
Ren G-D, C-X Yuan. 2005. Taxonomic study of the arboreal darking beetles from China Ainu Lewis 1894 (Coleoptera,Tenebrionidae). Acta Zootaxonomiga Sinica, 30(1): 98-103.
Triplehorn, C. A. 1985. A remarkable example of sexual dimorphism in Strongylium (Coleoptera: Tenebrionidae). Coleopterists-Bulletin, 39(1): 25-27.
Triplehorn, C. A., T. J. Spilman. 1973. A review of Strongylium of America North of Mexico, with descriptions of two new species (Coleptera,Tenebrionidae).Transactions American Entomological Society, 99(1): 1-27, figs. 1-25.
Tschinkel, W. R. 1975a. A comparative study of the chemical defensive system secretion of Tenebrionid beetles Ⅰ.chemistry of the secretions. J. Insect Physiol., 21: 753-783.
Tschinkel, W. R. 1975b. A comparative study of the chemical defensive system of Tenebrionid beetles Ⅱ. defensive behavior and ancillary features. Annals of the Entomological Society of America, 68(3): 439-453.
Tschinkel, W. R. 1975c. A comparative study of the chemical defensive system of Tenebrionid beetles Ⅲ. Morphology of the glands. Joun. Morph., 145: 355-370.
Tschinkel, W. R., J. T. Doyen. 1980. Comparative anatomy of the defensive glands, ovipositors and female genital tubes of Tenebrionid beetles (Coleoptera). Int. J. Insect Morphol. & Embryol.,9:321-386.
Watt,J. C. 1974. A revised subfamily classification of Tenebrionidae (Coleoptera). New Zealand Jour.Zool., 1:381-452.
Wu Chenfu F. 1936. Catalogus Insectorum Sinensium Ⅲ,Peking, 601-648.
Yuan C-X, G-D Ren. 2005. Study of the arboreal darkling beetles from China Strongylium Kirby 1818(Coleoptera,Tenebrionidae). Acta Zootaxonomiga Sinica, 30(2): 399-406.
李鸿兴.1997.鞘翅目.拟步甲科.杨星科主编.1997.长江三峡库区昆虫.重庆:重庆出版社,693.
刘晓丽.2004.部分拟步甲线粒体16S rDNA和18S rDNA基因序列与分子系统学研究[研究生毕业论文].河北大学.
任国栋,杨秀娟,印红.2002.基于防御腺的中国琵甲族Blaptini系统发育分析(鞘翅目:拟步甲科).昆虫学报,45(6):805-811.
任国栋,于有志.1999.中国荒漠半荒漠地区拟步甲分类研究.保定:河北大学出版社.
任国栋.1998.鞘翅目.拟步甲科.吴鸿主编.1998.龙王山昆虫.北京:中国林业出版社,111.
任国栋.2002.拟步甲科.黄邦侃主编.2002.福建昆虫志.第六卷.福州:福建科学技术出版社,137-170.
杨星科.1999.鞘翅目.郑乐怡,归鸿主编.1999.昆虫分类.南京:南京师范大学出版社,573.
叶晔,蒋鹏,任国栋.2003.31属拟步甲的防御腺特征与系统发育关系初析.河北大学学报(自然科学版),23(3):279-292.
于洋.2005.拟步甲成虫口器比较形态学研究(一).河北大学学报(自然科学版),25(3).(发表中).
张荣祖.1998.“中国动物地理区划”的再修订.动物分类学报,23(增刊):207-222.
张荣祖.1999.中国动物地理.北京:科学出版社.
赵养昌.1963.中国经济昆虫志.第四册.鞘翅目.伪步行虫科.北京:科学出版社.
周红章.2002.关于亚种下名称的可用性问题:实例与分析.动物分类学报,27(4):659-664.
周红章.2003.种级异物同名处理及属名变动影响分析.动物分类学报,28(1):5-12.
Ando, K. 2003. A new Strongylium species (Coleoptera:Tenebrionidae) occurring in Japan. Ent. Rev. Japan, 58(1):79-81, figs.1-2.
Arnett, R. H. Jr. 1968. The beetles of the United States (A manual for identification). Catholic University of America Press, Washington, D. C., 1112.
Bouchard, P. 2002. Phylogenetic revision of the flightless Australian genus Apterotheca Gebien (Coleoptera:Tenebrionidae: Coelometopinae). Invertebrate systematics,16: 449-554.
Bouchard, P., W. E. Steiner Jr. 2004. First descriptions of Coelometopini pupae (Coleoptera: Tenebrionidae) from Australia, Southeast Asia and the Pacific region, with comments on phylogenetie relationships and antipredator adaptations. Systematic Entomology, 29:101-102.
Chujo, M. 1968a. A check list of the Formosa Tenebrionidae (Coleoptera) Ⅱ. Kontyu, 36(1): 15-17, fig. 1.
Chujo, M. 1968b. A check list of the Formosa Tenebrionidae (Coleoptera) Ⅲ. Kontyu, 36(4): 397.
Chujo, M. 1978. Tenebrionidae of the Nansei Islands Ⅲ (Coleoptera). Esakia, (11): 78-80.
Chujo, M. 1979. Tenebrionidae of the Nansei Islands Ⅴ (Coleoptera). Esakia, (14): 54-56.
Chujo, M. 1998. Tenebrionidae (except genera Platydema Ischnodactylus, tribe Cnodalonini). In: Kurosawa, Y., Hisamatsu, S., Sasaji, H. 1998. the Coleoptera of Japan in color Vol. Ⅲ. Japan: Hoikusha Publishing Co., Ltd., 338-339.
Chujo, M., C. E. Lee. 1992. Tenebrionidae from Chejudo Island, Korea (Insecta, Coleoptera). Esakia, 32: 43-44, fig. 11.
Chujo, M., C. E. Lee. 1993. Korean Tenebrionidae (Insecta, Coleoptera). Esakia, 33:120. fig. 12,
Doyen, J. T. 1972. Familial and subfamilial classification of the Tenebrionidea (Coleoptera) and a revised generic classification of the Coniontini (Tentyriidae).Quaesitiones. Ent., 8(4): 357-367.
Doyen J. T. 1982. Phenetic and cladistie relationships among Tenebrionid beetles (Coleoptera). Syst. Entomol., 7(2): 127-183.
Doyen J. T. 1989. Reeonstitutions of Coelometopini Tenebrionini and related tribes of America north of Colombia (Coleoptera:Tenebrionidae). Journal of the New York Entomological Society, 97(3): 300-301, figs. 25-26.
Doyen J. T., J. F. Lawrence. 1979. Relationships and higher classification of some Tenebrionidae and Zopheridae (Coleoptera). Syst. Entomol., 4(4): 333-377.
Doyen J. T., R. E. Somerby. 1974. Phenetic similarity and mullerian mimicry among darking ground beetles(Coleoptera:Tenebrionidae).The Canadian Entomologist, 106: 771-772.
Fairmaire, L. 1891. Comptes-rendus des seances. Annales de la Societe Entomologique de Belgique, T., 35: 212-214.
Fairmaire, L. 1893. Coleopteres du Haut Tonkin. Annales de la Societe Entomologique de Belgique.,37: 319-321.
Fairmaire, L. 1893. Heteromeres. Annales de la Societe Entomologique de Belgique, 37:319-320.
Fairmaire, L. 1896. Note XII Coleopteres de L'inde Boreale, China et Malaisie. Notes from the Leyden Museum, 18:112-115.
Fairmaire, L. 1902. Descriotions de Coleopteres recueillis en Chine par M. de Latouche. Bulletin de la Societe Entomologique de France, 318.
Fairmaire, L. 1903. Descriptions de Quelques Heteromeres. Annales de la Societe Entomologique de Belgique, 47: 18-20.
Fairmaire, L., 1893. Note sur quelques Coleopteres des environs de Lang-Song. Annales de la Societe Entomologique de Belgique, 37: 299.
Gebien, H. 1913a. Die Tenebrionidae der Philippinen. The Philippine Journal of Science, 8(6):431-432.
Gebien, H. 1913b. H. Sauter's Formosa-A usbeute. Tenebrionidae (Coleoptera), Archiv fur Naturgeschichte, 79(A9): 44-54, figs. 1, 3, 4.
Gebien, H. 1944. Katalog der Tenebrioniden (Coleoptera) Ⅳ. Mitteilungen der Munchener Entomologischen Gesellschaft, 34:514(859)-555(899).
Imasaka, S. 1985. On the distributions of Strongylium marseuli Lewis and Strongylium cultellatum Maklin. Gekkan-Mushi, 172: 23-25.
Kaszab, Z. 1941. Tenebrioniden aus Formosa (Coleoptera). Stettiner Entomologische Zeitung, 102:71-72.
Kaszab, Z. 1954. Tenebrioniden Collected by J. Klapperich in Fukien. Ann. hist.-nat. Mus. Hung. (N. S.), 5: 247-264, figs.
Kaszab, Z. 1957. Neue Tenebrioniden (Coleoptera) aus Fukien. Bonn. Zool. Beitr., Bonn, 8(1): 56-63.
Kaszab, Z. 1964. Uber die Tenebrioniden einiger Japanischen Inseln (Ⅱ) (Coleoptera). Ent. Rev. Japan, Osaka,17: 8-10.
Kaszab, Z. 1977. Die Tenebrioniden des Papuanischen Gebietes Ⅰ. Strongyliini (Coleoptera: Tenebrionidae). Pacific Insects Monograph, 33: 1-219.
Kaszab, Z. 1980. Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici. 72:181.
Kulzer, H. 1966. Australische und Papuanische Strongyliini 29. Beitrag zur Kenntnis der Tenebrioniden. Ent. Arb. Mus. Frey, 338-353.
Kwon, Y. J., Y. S. Choi. 1986: Check list of Family Tenebrionidae from Korea. Insecta Koreana, 6:112-113
Lewis, G. 1894. On the Tenebrionidae of Japan. Annals & Magazine Natural History, 6(13): 479-483.
Maklin, F. W. 1864. Monographie der Gattung Strongylium Kirby, Lacordaire und der damit zunachst verwandten Formen. 518 pp., 4 pls. Finnlandischen Wissenschaftlichen Gesellschsft, Helsingfors.
Masumoto, K. 1981a. Tenebrionidae of Formosa (1). Elytra, 8(2): 37-53, figs. 1-20.
Masumoto, K. 1981b. Tenebrionidae of Formosa (2). Elytra, 9(1): 15-18, figs. 1-2.
Masumoto, K. 1981c. Tenebrionidae of Formosa (3). Elytra, 9(2): 95, 97, figs. 13-14.
Masumoto, K. 1981d. Notes and descriptions of Japanese Tenebrionidae (Ⅰ). Ent. Rev. Japan, 35:29-31, figs. 1-2.
Masumoto, K. 1982. Tenebrionidae of Formosa (5). Elytra, 10(2): 53-56, figs. 1, 1a, b, c, 2, 2a, b, c.
Masumoto, K. 1992. Tenebrionidae of East Asia (Ⅷ), Three new species of the genera Campsiomorpha (Cnodalonini) and Strongylium (Strongyliini) from Taiwan. Elytra, 20(1): 92-94, figs. 3, 4, 9-10.
Masumoto,K.1995.比和町立自然科学博物馆所藏台湾产科甲虫.(安藤+).比和科学 博物馆研究报告,33:17-22.
Masumoto, K. 1996a. New Tenebrionid beetles of the tribes Strongyliini, Misolampini and Adeliini (Coleoptera) from Northern Vietnam. Bull. Natn. Sci. Mus., Tokyo, (A), 22: 33-43.
Masumoto, K. 1996b. Replacement of the preoccupied name of a Strongylium species, Ent. Rev. Japan, 51: 14.
Masumoto, K. 1996c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅰ. Six new Strongylium species from Thailand, Laos and Taiwan, together with a new replacement name. Elytra, Tokyo, 24(1): 131-140, figs. 1-8.
Masumoto, K. 1996d. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅱ. New Strongylium species from Northern Thailand (Part 1). Elytra, Tokyo, 24(2): 337-366, figs. 1-20.
Masumoto, K. 1997a. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅲ. New Strongylium species from Southern India, preserved in the collections of the Museum National d'Histoire Naturelle, Paris and the Natural History Museum, London. Elytra, Tokyo, 25(1): 45-65, figs. 1-16.
Masumoto, K. 1997b. New records of Strongylium (Coleoptera,Tenebrionidae) from Northern Thailand. Elytra, Tokyo, 25(2): 65-66.
Masumoto, K. 1997c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅳ. Three species-groups of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 25(2): 279-309, figs. 1-56.
Masumoto, K. 1997d. New synonymy in the genus Strongylium (Coleoptera, Tenebrionidae, Strongyliini) described by Fairmaire and Pic. Elytra, Tokyo, 25(2): 309-310.
Masumoto, K. 1998a. New synonymy in the genus Strongylium (Coleoptera, Tenebrionidae, Strongyliini). Elytra, Tokyo, 26(1):200.
Masumoto, K. 1998b. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) V. Twenty new species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 26(1): 173-200, figs. 1-56.
Masumoto, K. 1998c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) Ⅵ. New Strongylium species from Northern Thailand (Part 2). Elytra, Tokyo, 26(2): 403-432, figs. 1-49.
Masumoto, K. 1999a. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅶ. Brachypterous strongyliines. Elytra, Tokyo, 27(1): 113-125, figs. 1-23.
Masumoto, K. 1999b. The male of Strongylium nanfangum (Coleoptera,Tenebrionidae). Elytra, Tokyo, 27(1):127-129.
Masumoto, K. 1999c. Study of Asian Strongyliini (Coleoptera,Tenebrionidae)Ⅷ.Ten new species of the genus Stongylium.Elytra,Tokyo, 27(2): 335-352, figs.1-31.
Masumoto, K. 2000a. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) IX. Hairy Strongylium species from Southeast Asia (part Ⅰ. Species-group of Strongylium gibbosulum). Elytra, Tokyo, 28(1): 163-172, figs. 1-17.
Masumoto, K. 2000b. Study of Asian Strongyliini (Coleoptera,Tenebrionidae) X. Ten new species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 28(2): 391-407, figs. 1-31.
Masumoto, K. 2001. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) XI. Ten new small-sized species of the genus Strongylium from Southeast Asia. Elytra, Tokyo, 29(2): 401-418, figs. 1-30.
Masumoto, K. 2002. A new Strongylium (Coleoptera,Tenebrionidae, Strongyliini) from Borneo. Elytra, Tokyo, 30(2): 335-338. figs. 1-3.
Masumoto, K. 2003. Study of Asian Strongyliini (Coleoptera, Tenebrionidae) Ⅻ. New Strongylium species from Thailand. Elytra, 31(1): 65-86.
Medvedev, G. S. 1977. Taxonomical study of antennal sensillia in Tenebrionidae (Coleoptera: Tenebrionidae). Trudy Vses. Ent. Obshch, 58: 61-86, illust. [in Russian]
Miwa, Y. 1931. A systematic catalogue of Formosan Coleoptera. Rept. 55 Dept. Agr. Gov. Res. Inst. Formosa, 166.
Miwa, Y. 1939. Descriptions on four new species belonging to the family Tenebrionidae from Formosa. Zoological Magazine, Tokyo, 51(7): 414-415, figs. 3-4.
Nakane, T. 1968. New or little-known Coleoptera from Japan and its adjacent regions 27. Fragmenta Coleopterol., 21: 83-85.
Nomura, S. 1963: Some new species of the Coleoptera from Loochoo Is. and its adjacent regions, The Entpmological Review of Japan, 15(2): 37-40; P1.5, figs. 1-4.
Nomura, S. 1964: Some new species of the Coleoptera from Loochoo Is. and its Adjacent Regions, Ⅱ. The Entpmological Review of Japan, 17(2): 47-49. P1.3, fig. 4.
Pic, M. 1916. Especes et varietes nouvelles. Melanges Exotico-Entomologiques, 19: 12-13.
Pic, M. 1917. Coleopteres exotiques en partie nouveaux. L'Echange, Revue Linneenne, 33: 12.
Pic, M. 1928. Cinquantr-et-unieme fascicule. Melanges Exotico Entomologiques, 51: 23-24.
Pic, M. 1936. Nouveautes diverses. Melanges Exotico Entomologiques, 68: 19-20.
Pic, M. 1940. Diagnoses de Coleopteres exotiques. L'Echange, Revue Linneenne, 56:13.
Reitter, E. 1904. Bestimmungs-tabelle Tenebrioniden-unterfanilien: Lachnogyini, Akidini, Opatrini und Trachyscelini aus Europa und den angrenzenden Landern. Verh. Nat.-forsch. Ver. Brunn,12:25-189.
Ren G-D, C-X Yuan. 2005. Taxonomic study of the arboreal darking beetles from China Ainu Lewis 1894 (Coleoptera,Tenebrionidae). Acta Zootaxonomiga Sinica, 30(1): 98-103.
Triplehorn, C. A. 1985. A remarkable example of sexual dimorphism in Strongylium (Coleoptera: Tenebrionidae). Coleopterists-Bulletin, 39(1): 25-27.
Triplehorn, C. A., T. J. Spilman. 1973. A review of Strongylium of America North of Mexico, with descriptions of two new species (Coleptera,Tenebrionidae).Transactions American Entomological Society, 99(1): 1-27, figs. 1-25.
Tschinkel, W. R. 1975a. A comparative study of the chemical defensive system secretion of Tenebrionid beetles Ⅰ.chemistry of the secretions. J. Insect Physiol., 21: 753-783.
Tschinkel, W. R. 1975b. A comparative study of the chemical defensive system of Tenebrionid beetles Ⅱ. defensive behavior and ancillary features. Annals of the Entomological Society of America, 68(3): 439-453.
Tschinkel, W. R. 1975c. A comparative study of the chemical defensive system of Tenebrionid beetles Ⅲ. Morphology of the glands. Joun. Morph., 145: 355-370.
Tschinkel, W. R., J. T. Doyen. 1980. Comparative anatomy of the defensive glands, ovipositors and female genital tubes of Tenebrionid beetles (Coleoptera). Int. J. Insect Morphol. & Embryol.,9:321-386.
Watt,J. C. 1974. A revised subfamily classification of Tenebrionidae (Coleoptera). New Zealand Jour.Zool., 1:381-452.
Wu Chenfu F. 1936. Catalogus Insectorum Sinensium Ⅲ,Peking, 601-648.
Yuan C-X, G-D Ren. 2005. Study of the arboreal darkling beetles from China Strongylium Kirby 1818(Coleoptera,Tenebrionidae). Acta Zootaxonomiga Sinica, 30(2): 399-406.
© 2004-2018 中国地质图书馆版权所有 京ICP备05064691号 京公网安备11010802017129号 地址：北京市海淀区学院路29号 邮编：100083 电话：办公室：(+86 10)66554848；文献借阅、咨询服务、科技查新：66554700 |