中国钩瓣叶蜂属系统分类研究
摘要
本文系统地回顾了钩瓣叶蜂属Macrophya Dahlbom分类研究历史，介绍了钩瓣叶蜂属的外部形态、分类特征、生物学习性；在简要分析了钩瓣叶蜂属地理分布状况的基础上，初步探讨了钩瓣叶蜂属的分布格局和可能的成因，结果表明，喜马拉雅山及横断山的东南部很可能是钩瓣叶蜂属的现代分布中心和分化中心；本文还深入研究了中国钩瓣叶蜂属系统分类，编制了中国钩瓣叶蜂属分种检索表，依据标本及文献共记述110种，其中包括2个中国新记录种，41个新种：每种都有完备的引证、形态描述、鉴别特征、分布及检视标本记录。绝大多数种类均附有雌雄外生殖器特征图。新种的模式标本保存于中南林学院昆虫资源研究所昆虫模式标本室。中国新记录种和新种名单如下：
新种：
吉林钩瓣叶蜂 M. jilinensis Chen et Wei, sp. nov.
朱氏钩瓣叶蜂 M. zhui Chen et Wei, sp. nov.
西藏钩瓣叶蜂 M. tibetana Chen et Wei, sp. n
白跗钩瓣叶蜂 M. leucotarsalina Chen et Wei, sp. nov.
缩唇钩瓣叶蜂 M. breviclypea Chen et Wei, sp. n
神龙钩瓣叶蜂 M. shennongjiana Chen et Wei, sp. nov.
锐唇钩瓣叶蜂 M. acuminiclypea Chen et Wei, sp. nov.
粗纹钩瓣叶蜂 M. megasculpturalis Chen et Wei, sp. nov.
细瓣钩瓣叶蜂 M. parviserrula Chen et Wei, sp. nov.
黄斑钩瓣叶蜂井冈亚种 M. fiavomaculata jinggangensis Chen et Wei, sp. nov.
宝石钩瓣叶蜂 M. xanthosoma Chen et Wei, sp. nov.
黑脊钩瓣叶蜂 M. histrio nigrihistrio Chen et Wei, sp. nov.
低额钩瓣叶蜂 M. frontalis Chen et Wei, sp. nov.
异碟钩瓣叶蜂 M. allominutifossa Chen et Wei, sp. nov.
三斑钩瓣叶蜂 M. trimicralba Chen et Wei, sp. nov.
长钩钩瓣叶蜂 M. paracoxalis Chen et Wei, sp. nov.
卜氏钩瓣叶蜂 M. bui Chen et Wei, sp. nov.
九寨钩瓣叶蜂 M. fiuzhaina Chen et Wei, sp. n
淡痣钩瓣叶蜂 M. fulvostigmata Chen et Wei, sp. nov.
钟氏钩瓣叶蜂 M. zhongi Wei et Chen, sp. nov.
点斑钩瓣叶蜂 M. minutiluna Chen et Wei, sp. nov.
五斑钩瓣叶蜂 M. pentanalia Chen et Wei, sp. nov.
江氏钩瓣叶蜂 M. jiangi Chen et Wei, sp. nov.
糙额钩瓣叶蜂 M. opacifrontalis Chen et Wei, sp. nov.
长白钩瓣叶蜂 M. changbaina Chen et Wei, sp. nov.
任氏钩瓣叶蜂 M. reni Chen et Wei, sp. n
雷氏钩瓣叶蜂 M. leyii Chen et Wei, sp. nov.
暗唇钩瓣叶蜂 M. melanoclypea Chen et Wei, sp. nov.
黄氏钩瓣叶蜂 M. huangi Chen et Wei, sp. nov.
6 中国钧肘叶峰周系统分类研究
白转钩瓣叶蜂 M alboDDeaa’ztera Chen et We，SI）．nov．
横脊钩瓣叶蜂 M呻ena Chen et We Sp．nov
凹颜钩瓣叶蜂 M epressina Chen et Wei，Sp．nov
斑瓣钩瓣叶蜂M。。lipe。mso伽etwe，SI）．nov．
肿附钩瓣叶蜂M C——加一加a ChCh一to乃，叩．lOV．
木氏钩瓣叶蜂 M mui ChChen eto厄，Sp．n
缩臀钩瓣叶蜂Moo。0止砌V厄et戳en，Sp．nov．
平刃钩瓣叶蜂 M dectose。b Chen et Wei，sp．nov．
白唇钩瓣叶蜂 M albolabrtana Chen et Wei，sp nov，
甘肃钩瓣叶蜂 M g。us。iCQ Chgh et Wei，Sp．DOV．
拟鞘钩瓣叶蜂 M imitato沾幻 Chen etwe，叩．nov．
结窝钩瓣叶蜂／no颂viata Chen et Wei，sp．nov．
中国新记录：
平盾钩瓣叶蜂 M planatQ MOcsaryl 909
密鞘钩瓣叶蜂MzmjmjorhkeUch，1937
The research histories of the genus Macrophya Dahlbom are briefly reviewed. The morphology, taxonomic and biological characters of the genus are outlined. Based on the analysis of the generic and specific distribution of Macrophya Dahlbom the distribution pattern and the possible origination of the genus is discussed. It is seemed that east-southern region of Mt. Himalayas and Mt. Hengduan with neighbouring area is probably the derivation center of Macrophya. The taxonomy of Macrophya from China is also studied in detail in the thesis. Key to known species of the genus from China are given. Totally 110 species from China are described. Among them 41 species are new to science and 2 species are new records in China. To each species full incitations, morphological description, diagnostic characters, distribution, specimens examined are provided, as well as female and male genitalia illustrations of almost all species. Type specimens are deposited in the bisect Collection of Institute of Insect Resources of Central South Forestry University, Zhuzhou, China. The new species and new recorded species from China are as folio wings:
New species:
M. jilinensis Chen et Wei, sp. nov.
M. zhui Chen et Wei, sp. nov.
M. tibetana Chen et Wei, sp. nov.
M. leucotarsalina Chen et Wei, sp. nov.
M. breviclypea Chen et Wei, sp. nov.
M. shennongjiana Chen et Wei, sp. nov.
M. acuminiclypea Chen et Wei, sp. nov.
M. megasculpturalis Chen et Wei, sp. nov.
M. parviserrula Chen et Wei, sp. nov.
M. flavomaculata jinggangensis Chen et Wei, sp. nov.
M. xanthosoma Chen et Wei, sp. nov.
M. histrio nigrihistrio Chen et Wei, sp. nov.
M. frontalis Chen et Wei, sp. nov.
M. allominutifossa Chen et Wei, sp. nov.
M. trimicralba Chen et Wei, sp. nov.
M. paracoxalis Chen et Wei, sp. nov.
M. bui Chen et Wei, sp. n
M. jiuzhaina Chen et Wei, sp. nov.
M. fulvostigmata Chen et Wei, sp. nov.
M. zhongi Chen et Wei, sp. nov.
M. minutiluna Chen et Wei, sp. nov.
M. pentanalia Chen et Wei, sp. nov. %
M.jiangi Chen et Wei, sp. nov.
M. opacifrontalis Chen et Wei, sp. nov.
M. changbaina Chen et Wei, Sp. nov.
M. rent Chen et Wei, sp. nov.
M. Leyii Chen et Wei, sp. nov.
M. melanoclypea Chen e/ Wei, sp. nov.
M. huangi Chen ef Wei, sp. nov.
M. albotrochantera Chen et Wei, sp. nov.
M. tripidona Chen e/ Wei, sp. nov.
M. depressina Chen et Wei, sp. nov.
M. maculipennis Chen e/ Wei, sp. nov.
M. crassitarsalina Chen e/ Wei, sp. nov.
M. mui Chen et Wei, sp. nov.
M. constrictila Chen e/ Wei, sp. nov.
M. flactoserrula Chen et Wei, sp. nov.
M albolabriana Chen e/ Wei, sp. nov.
M. gansuica Chen e/ Wei, sp. nov.
M. imitatoides Chen et Wei, sp. nov.
M. nodifoviata Chen ef Wei, sp. nov.
Species firstly recorded from China:
M. planata Mocsary, 1909 M. imitator Takeuchi, 1937
新种：
吉林钩瓣叶蜂 M. jilinensis Chen et Wei, sp. nov.
朱氏钩瓣叶蜂 M. zhui Chen et Wei, sp. nov.
西藏钩瓣叶蜂 M. tibetana Chen et Wei, sp. n
白跗钩瓣叶蜂 M. leucotarsalina Chen et Wei, sp. nov.
缩唇钩瓣叶蜂 M. breviclypea Chen et Wei, sp. n
神龙钩瓣叶蜂 M. shennongjiana Chen et Wei, sp. nov.
锐唇钩瓣叶蜂 M. acuminiclypea Chen et Wei, sp. nov.
粗纹钩瓣叶蜂 M. megasculpturalis Chen et Wei, sp. nov.
细瓣钩瓣叶蜂 M. parviserrula Chen et Wei, sp. nov.
黄斑钩瓣叶蜂井冈亚种 M. fiavomaculata jinggangensis Chen et Wei, sp. nov.
宝石钩瓣叶蜂 M. xanthosoma Chen et Wei, sp. nov.
黑脊钩瓣叶蜂 M. histrio nigrihistrio Chen et Wei, sp. nov.
低额钩瓣叶蜂 M. frontalis Chen et Wei, sp. nov.
异碟钩瓣叶蜂 M. allominutifossa Chen et Wei, sp. nov.
三斑钩瓣叶蜂 M. trimicralba Chen et Wei, sp. nov.
长钩钩瓣叶蜂 M. paracoxalis Chen et Wei, sp. nov.
卜氏钩瓣叶蜂 M. bui Chen et Wei, sp. nov.
九寨钩瓣叶蜂 M. fiuzhaina Chen et Wei, sp. n
淡痣钩瓣叶蜂 M. fulvostigmata Chen et Wei, sp. nov.
钟氏钩瓣叶蜂 M. zhongi Wei et Chen, sp. nov.
点斑钩瓣叶蜂 M. minutiluna Chen et Wei, sp. nov.
五斑钩瓣叶蜂 M. pentanalia Chen et Wei, sp. nov.
江氏钩瓣叶蜂 M. jiangi Chen et Wei, sp. nov.
糙额钩瓣叶蜂 M. opacifrontalis Chen et Wei, sp. nov.
长白钩瓣叶蜂 M. changbaina Chen et Wei, sp. nov.
任氏钩瓣叶蜂 M. reni Chen et Wei, sp. n
雷氏钩瓣叶蜂 M. leyii Chen et Wei, sp. nov.
暗唇钩瓣叶蜂 M. melanoclypea Chen et Wei, sp. nov.
黄氏钩瓣叶蜂 M. huangi Chen et Wei, sp. nov.
6 中国钧肘叶峰周系统分类研究
白转钩瓣叶蜂 M alboDDeaa’ztera Chen et We，SI）．nov．
横脊钩瓣叶蜂 M呻ena Chen et We Sp．nov
凹颜钩瓣叶蜂 M epressina Chen et Wei，Sp．nov
斑瓣钩瓣叶蜂M。。lipe。mso伽etwe，SI）．nov．
肿附钩瓣叶蜂M C——加一加a ChCh一to乃，叩．lOV．
木氏钩瓣叶蜂 M mui ChChen eto厄，Sp．n
缩臀钩瓣叶蜂Moo。0止砌V厄et戳en，Sp．nov．
平刃钩瓣叶蜂 M dectose。b Chen et Wei，sp．nov．
白唇钩瓣叶蜂 M albolabrtana Chen et Wei，sp nov，
甘肃钩瓣叶蜂 M g。us。iCQ Chgh et Wei，Sp．DOV．
拟鞘钩瓣叶蜂 M imitato沾幻 Chen etwe，叩．nov．
结窝钩瓣叶蜂／no颂viata Chen et Wei，sp．nov．
中国新记录：
平盾钩瓣叶蜂 M planatQ MOcsaryl 909
密鞘钩瓣叶蜂MzmjmjorhkeUch，1937
The research histories of the genus Macrophya Dahlbom are briefly reviewed. The morphology, taxonomic and biological characters of the genus are outlined. Based on the analysis of the generic and specific distribution of Macrophya Dahlbom the distribution pattern and the possible origination of the genus is discussed. It is seemed that east-southern region of Mt. Himalayas and Mt. Hengduan with neighbouring area is probably the derivation center of Macrophya. The taxonomy of Macrophya from China is also studied in detail in the thesis. Key to known species of the genus from China are given. Totally 110 species from China are described. Among them 41 species are new to science and 2 species are new records in China. To each species full incitations, morphological description, diagnostic characters, distribution, specimens examined are provided, as well as female and male genitalia illustrations of almost all species. Type specimens are deposited in the bisect Collection of Institute of Insect Resources of Central South Forestry University, Zhuzhou, China. The new species and new recorded species from China are as folio wings:
New species:
M. jilinensis Chen et Wei, sp. nov.
M. zhui Chen et Wei, sp. nov.
M. tibetana Chen et Wei, sp. nov.
M. leucotarsalina Chen et Wei, sp. nov.
M. breviclypea Chen et Wei, sp. nov.
M. shennongjiana Chen et Wei, sp. nov.
M. acuminiclypea Chen et Wei, sp. nov.
M. megasculpturalis Chen et Wei, sp. nov.
M. parviserrula Chen et Wei, sp. nov.
M. flavomaculata jinggangensis Chen et Wei, sp. nov.
M. xanthosoma Chen et Wei, sp. nov.
M. histrio nigrihistrio Chen et Wei, sp. nov.
M. frontalis Chen et Wei, sp. nov.
M. allominutifossa Chen et Wei, sp. nov.
M. trimicralba Chen et Wei, sp. nov.
M. paracoxalis Chen et Wei, sp. nov.
M. bui Chen et Wei, sp. n
M. jiuzhaina Chen et Wei, sp. nov.
M. fulvostigmata Chen et Wei, sp. nov.
M. zhongi Chen et Wei, sp. nov.
M. minutiluna Chen et Wei, sp. nov.
M. pentanalia Chen et Wei, sp. nov. %
M.jiangi Chen et Wei, sp. nov.
M. opacifrontalis Chen et Wei, sp. nov.
M. changbaina Chen et Wei, Sp. nov.
M. rent Chen et Wei, sp. nov.
M. Leyii Chen et Wei, sp. nov.
M. melanoclypea Chen e/ Wei, sp. nov.
M. huangi Chen ef Wei, sp. nov.
M. albotrochantera Chen et Wei, sp. nov.
M. tripidona Chen e/ Wei, sp. nov.
M. depressina Chen et Wei, sp. nov.
M. maculipennis Chen e/ Wei, sp. nov.
M. crassitarsalina Chen e/ Wei, sp. nov.
M. mui Chen et Wei, sp. nov.
M. constrictila Chen e/ Wei, sp. nov.
M. flactoserrula Chen et Wei, sp. nov.
M albolabriana Chen e/ Wei, sp. nov.
M. gansuica Chen e/ Wei, sp. nov.
M. imitatoides Chen et Wei, sp. nov.
M. nodifoviata Chen ef Wei, sp. nov.
Species firstly recorded from China:
M. planata Mocsary, 1909 M. imitator Takeuchi, 1937
引文
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62． Rohwer S A. 1916． Chalastogastra (Hym.). Suppl. Ent., 5: 81-113．
63． Rohwer S A. 1921． Notes on sawflies, with descriptions of new genera and species. Proc. U. S. Nat. Mus., 59:83-109．
64． Rohwer S A. 1925． Sawflies from the Maritime Province of Siberia. Proc. U. S. Nat. Mus., 68: 1-12．
65． Rohwer S A. 1934． On the Tenthredinoidea of the Florissant Shalis. Bulletin American Museum of Natural History, 24: 521-530
66． Ross H H. 1933． The Description and life history of a new sawfly Sterictiphora apios (Argidae: Hymen. ). Proc. Ent. Soc. Washington., 35(2) : 13-19
67． Ross H H. 1936． The ancestry and wing venation of Hymenoptera. Ann. Ent. Soc. Amer., 29: 99-111．
68． Ross H H. 1937． A generic classification of the Nearctic sawflies (Hymenoptera Symphyta). University of Illinois Bulletin, Vol. 34, N 94: Illinois Biol Monographs, Vol. XV, N 2, Urbana, Illinois, pp. 1-173
69． Ross H H. 1951． Suborder Symphyta (Chalastogastra).-In: Muesebeck of al., Argidae. Monographs, No. 2, Washington: 4-89
70． Saini M S. & Dhillon S S. 1981． Evolutionary changes in the symphytancranium pertaining to epicranial and pauoccipital sutures ( Hymenoptera). J. Kansas Ent. Soc., 4: 458-464．
71． Saini M S. 1996． Taxonomic Revision of genus Macrophya Dahlbom from India. Dtsch. Ent. Z., 43: 129-154
72． Smith D R. 1979． Catalogue of Hymenoptera in America N. Mexico. Volume 1: Symphyta and Apocrita. Pp. 26-29
73． Smith D R. Gibson G A. P. 1984． Filacus, a new Genus for four species of sawflies previously placed in Macrophya or Zaschizonyx (Hym. : Tenthredinidae). Pan Pac. Ent., 60(2) 101-113
74． Smith D R. and Barrows E. 1987． Sawflies (Hymenoptera: Symphyta) in urbanenvironments in the Washington, D. C. area. Proc. Ent. Soc. Wash., 89: 147-156
75． Smith D R. and Middlekauff W W. 1987． Suborder Symphyta.. in Stehr, F. W., ed., Immature insects. Kendall Hunt Publishing Company, DubuSmith, D. R. ue, Iowa.: 754
76． Smith D R. 1988． A synopsis of the sawflies of America south of the United States: introduction, Xyelidae, Pamphiliidae, Cimbicidae, Diprionidae, Xipbydriidae, Siricidae, Orussidae, Cephidae. Systematic Entomology, 13: 205-261
77． Taeger A. 1989． Die Gattuang Macrophya D. In der DDR (Insecta, Hym. Symphyta: Tenthredinidae). :57-69
78． Takeuchi K. 1927． Some Chalastogastra from Corea. Trans. Nat. Hist. Soc. Formosa, 17: 378-387
79． Takeuchi K. 1931． Some sawflies from the Island of Shikoku in the southern Kuriles. Trans. Kansai Ent. Soc., No.2． : 30-34．
80． Takeuchi K. 1937． A study on the Japanese species of the genus Macrophya Dahlbom (Hymenoptera Tenthredinidae). Tenthredo, 1: 376-454．
81． Takeuchi K. 1938． A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (I). Tenthredo, II(2) : 173-229
82． Takeuchi K. 1938． Chinese Sawflies and wood wasps in the collection of the Musee Heude in Shanghai. Notes d'Ent. Chi., 5: 59-85．
83． Takeuchi K. 1939． A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire(II). Tenthredo, 2(4) : 393-439．
84． Takeuchi K. 1988． Symphyta of Japanese Empire (II). Tenthredo, II(4) : 82-105
85． Thomson C G 1871． Hymenoptera Scandinavia, Vol. 1． Phytophaga. Lund, Ohlsson: 342
86． Togashi. I. 1970． New genus and species of the Sterictiphorinae from Japan. Mushi., 44: 49-53
87． Togashi I. 1976． Sawflies (Hymenoptera: Tenthredinidae) of Taiwan collected by 124． Mr. N. Fukuhara in 1971． Kontyu, 44(4) : 477-479
88． Togashi I. 1988． Symphyta of Thailand (Insecta, Hymenoptera). Steenstrupia, 14: 101-119
89． Togashi I. 1990． Sawflies of Chejudo Island, Korea. Ent. Journ. Fukui. No. 7: 21-24．
90． Turner R E. 1919． On Indo-Chinese Hymenoptera collected by R.Vitalisde Salvaza. Ann. Mag. Nat. Hist., : 84-98．
91． Wei M. 1998． New species of Macrophya from Funiushan (Hymenoptera: Tenthredinidae). The Fauna and Taxonomy of Insects in Henan, Vol. 4: 152-161
92． Wei M, H Nie. 1999． New species of sawflies collected by Mr. Sheng and Ms. Sun from Henan province. The fauna and taxonomy of insect in Henan, Vol. 4: 160-161．
93． Wei M, H Nie. 1999． New species of Macrophya from south slope of Mt. Funiu. The fauna and taxonomy of insect in Henan, Vol. 4,: 104．
94． Wei M, H Nie. 1998． New species of Macrophya from Funiushan. The fauna and taxonomy of insect in Henan, Vol. 2． 152-161
95． Wei M, H Nie. 1997． Five new species of Macrophya from China. Entomotaxonomia, 19, Suppl.: 77-84
96． Wei M, H Nie. 1997． A new genus and eight new species of Tenthredinidae from Jiangxi. Entomotaxonomia., 19(1) : 70-73
97． Westwood J O. 1840． Introduction to the Modeem Classification of Insects, II, Longman, Orme, Brown. Green, and Longmans, London.
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100． Zhelochovtsev A H. 1934． Ueber die Typen der japanischen Tenthredinidae in W. Motschulski Zammiung. Arch. Mus. Zool. Univ. Moscou, 1: 147-149．
101． Zhelochovtsev A H. 1988． Order Hymenoptera. Suborder Symphyta in Opcedelitel nosekomych evropejskoj casti SSSR. ToM. 3． Hymenoptera part, 6 : 3-53
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58． Rohwer S A. 1910． Japanese sawflies in the collection of the United States National 95． Museum Proc. U. S. Nat. Mas., 39: 99-120．
59． Rohwer S A. 1911． The genotypes of sawflies and wood wasps, or the superfamily Tenthredinoidea. U. S. Bull. Entomol. Tech., Ser., 20: 69-109．
60． Rohwer S A. 1915． Some oriental sawflies in the Indian Museum. Rec. Ind. Mus., 11: 39-53．
61． Rohwer S A. 1915． The mating habits of some sawflies. Proc. Wash. Ent Soc., 17: 195-198．
62． Rohwer S A. 1916． Chalastogastra (Hym.). Suppl. Ent., 5: 81-113．
63． Rohwer S A. 1921． Notes on sawflies, with descriptions of new genera and species. Proc. U. S. Nat. Mus., 59:83-109．
64． Rohwer S A. 1925． Sawflies from the Maritime Province of Siberia. Proc. U. S. Nat. Mus., 68: 1-12．
65． Rohwer S A. 1934． On the Tenthredinoidea of the Florissant Shalis. Bulletin American Museum of Natural History, 24: 521-530
66． Ross H H. 1933． The Description and life history of a new sawfly Sterictiphora apios (Argidae: Hymen. ). Proc. Ent. Soc. Washington., 35(2) : 13-19
67． Ross H H. 1936． The ancestry and wing venation of Hymenoptera. Ann. Ent. Soc. Amer., 29: 99-111．
68． Ross H H. 1937． A generic classification of the Nearctic sawflies (Hymenoptera Symphyta). University of Illinois Bulletin, Vol. 34, N 94: Illinois Biol Monographs, Vol. XV, N 2, Urbana, Illinois, pp. 1-173
69． Ross H H. 1951． Suborder Symphyta (Chalastogastra).-In: Muesebeck of al.
70． Saini M S. & Dhillon S S. 1981． Evolutionary changes in the symphytancranium pertaining to epicranial and pauoccipital sutures ( Hymenoptera). J. Kansas Ent. Soc., 4: 458-464．
71． Saini M S. 1996． Taxonomic Revision of genus Macrophya Dahlbom from India. Dtsch. Ent. Z., 43: 129-154
72． Smith D R. 1979． Catalogue of Hymenoptera in America N. Mexico. Volume 1: Symphyta and Apocrita. Pp. 26-29
73． Smith D R. Gibson G A. P. 1984． Filacus, a new Genus for four species of sawflies previously placed in Macrophya or Zaschizonyx (Hym. : Tenthredinidae). Pan Pac. Ent., 60(2) 101-113
74． Smith D R. and Barrows E. 1987． Sawflies (Hymenoptera: Symphyta) in urbanenvironments in the Washington, D. C. area. Proc. Ent. Soc. Wash., 89: 147-156
75． Smith D R. and Middlekauff W W. 1987． Suborder Symphyta.. in Stehr, F. W., ed., Immature insects. Kendall Hunt Publishing Company, DubuSmith, D. R. ue, Iowa.: 754
76． Smith D R. 1988． A synopsis of the sawflies of America south of the United States: introduction, Xyelidae, Pamphiliidae, Cimbicidae, Diprionidae, Xipbydriidae, Siricidae, Orussidae, Cephidae. Systematic Entomology, 13: 205-261
77． Taeger A. 1989． Die Gattuang Macrophya D. In der DDR (Insecta, Hym. Symphyta: Tenthredinidae). :57-69
78． Takeuchi K. 1927． Some Chalastogastra from Corea. Trans. Nat. Hist. Soc. Formosa, 17: 378-387
79． Takeuchi K. 1931． Some sawflies from the Island of Shikoku in the southern Kuriles. Trans. Kansai Ent. Soc., No.2． : 30-34．
80． Takeuchi K. 1937． A study on the Japanese species of the genus Macrophya Dahlbom (Hymenoptera Tenthredinidae). Tenthredo, 1: 376-454．
81． Takeuchi K. 1938． A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (I). Tenthredo, II(2) : 173-229
82． Takeuchi K. 1938． Chinese Sawflies and wood wasps in the collection of the Musee Heude in Shanghai. Notes d'Ent. Chi., 5: 59-85．
83． Takeuchi K. 1939． A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire(II). Tenthredo, 2(4) : 393-439．
84． Takeuchi K. 1988． Symphyta of Japanese Empire (II). Tenthredo, II(4) : 82-105
85． Thomson C G 1871． Hymenoptera Scandinavia, Vol. 1． Phytophaga. Lund, Ohlsson: 342
86． Togashi. I. 1970． New genus and species of the Sterictiphorinae from Japan. Mushi., 44: 49-53
87． Togashi I. 1976． Sawflies (Hymenoptera: Tenthredinidae) of Taiwan collected by 124． Mr. N. Fukuhara in 1971． Kontyu, 44(4) : 477-479
88． Togashi I. 1988． Symphyta of Thailand (Insecta, Hymenoptera). Steenstrupia, 14: 101-119
89． Togashi I. 1990． Sawflies of Chejudo Island, Korea. Ent. Journ. Fukui. No. 7: 21-24．
90． Turner R E. 1919． On Indo-Chinese Hymenoptera collected by R.Vitalisde Salvaza. Ann. Mag. Nat. Hist., : 84-98．
91． Wei M. 1998． New species of Macrophya from Funiushan (Hymenoptera: Tenthredinidae). The Fauna and Taxonomy of Insects in Henan, Vol. 4: 152-161
92． Wei M, H Nie. 1999． New species of sawflies collected by Mr. Sheng and Ms. Sun from Henan province. The fauna and taxonomy of insect in Henan, Vol. 4: 160-161．
93． Wei M, H Nie. 1999． New species of Macrophya from south slope of Mt. Funiu. The fauna and taxonomy of insect in Henan, Vol. 4,: 104．
94． Wei M, H Nie. 1998． New species of Macrophya from Funiushan. The fauna and taxonomy of insect in Henan, Vol. 2． 152-161
95． Wei M, H Nie. 1997． Five new species of Macrophya from China. Entomotaxonomia, 19, Suppl.: 77-84
96． Wei M, H Nie. 1997． A new genus and eight new species of Tenthredinidae from Jiangxi. Entomotaxonomia., 19(1) : 70-73
97． Westwood J O. 1840． Introduction to the Modeem Classification of Insects, II, Longman, Orme, Brown. Green, and Longmans, London.
98． Wu Chenfu F. 1941． Order XXV. Hymenoptera. Wu, Catalogus Insectorum Sinensium. VI: 1-8
99． Yuasa H. 1922． A classification of the larvae of the Tenthredinoidea. Illinois Biol. Monographs, VII(4) : 172
100． Zhelochovtsev A H. 1934． Ueber die Typen der japanischen Tenthredinidae in W. Motschulski Zammiung. Arch. Mus. Zool. Univ. Moscou, 1: 147-149．
101． Zhelochovtsev A H. 1988． Order Hymenoptera. Suborder Symphyta in Opcedelitel nosekomych evropejskoj casti SSSR. ToM. 3． Hymenoptera part, 6 : 3-53
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