盐城保护区底栖动物及鸻鹬类群落研究
|
摘要
鸻鹬类是典型的湿地鸟类,是海岸和湖泊等自然生态系统中的一个非常重要的组成成分,其数量的多少是衡量湿地重要性的标准之一。中途停歇地的食物供应对鸻鹬类补充能量以便顺利完成迁徙并成功繁殖至关重要。盐城自然保护区是东亚-澳大利亚候鸟迁徙路线上鸻鹬类重要的中途停歇地,为了研究盐城自然保护区对鸻鹬类的支持作用,为鸻鹬类的保护提供科学依据,本课题选在射阳河口滩涂,研究该区内作为鸻鹬类食物的大型底栖动物的资源情况、分析潮间带底栖动物的生态位及互花米草对大型底栖动物的影响；继而对该区迁徙期、越冬期鸻鹬类的资源情况以及春、冬2季大型底栖动物和鸻鹬类的关系进行研究。主要结果如下：
1、研究地潮间带共有大型底栖动物16种,各生境的物种数不同。生境条件与潮间带大型底栖动物群落组成结构有一定的关系,同一生境大型底栖动物群落季节间差异不显著。焦河蓝蛤(Potamocorbula ustulata)和明细白樱蛤(Macoma praetexta)的生态位较宽,中国绿螂(Glaucomya chinensis)和四角蛤蜊(Mactra veneriformis)较窄。不同的物种间生态位重叠值差异较大。物种生态位宽度、物种之间的生态位重叠值与物种的时空分布和数量密切相关。
2、研究地滩涂共发现大型底栖动物22种。米草区和非米草区的物种组成不同,不同生境中的互花米草对底栖动物的影响不甚相同：潮间带互花米草的入侵降低了大型底栖动物的物种数、密度、生物量和多样性；潮沟米草的入侵提高了大型底栖动物的物种数、密度和多样性；而潮上带米草入侵对大型底栖动物的各指标影响不一。
3、研究地内共统计到鸻鹬类22种,这些种类在春季均能看到,秋季13种,冬季仅8种；黑腹滨鹬(Calidris alpina)是3个季节共同的优势种,春季、秋季和冬鸻鹬类的数量比值约为3：1：1.5；3个季节鸻鹬类的密度在生境间的差异均显著；3个季节,鸻鹬类物种数和Shannon-weinner多样性指数最多的均是潮沟,密度较高的是高潮带光滩、中潮带光滩和潮沟,数量最多的高潮带光滩和中潮带光滩；养殖塘也有一定数量的鸻鹬类分布,但密度较低,多以栖息为主；春季多数鸟类的迁徙高峰在4月中下旬,结束于5月中下旬,不同的类群迁徙高峰略有差异。
4、底栖动物密度较高的生境,鸻鹬类的密度也较高。春季,嘴长50mmm以上鸻鹬类的密度与甲壳类密度显著正相关,嘴长25mm以下鸻鹬类密度与双壳类密度及底栖动物总密度显著正相关,嘴长25-50mm鸻鹬类密度与任何一类底栖动物均未表现出明显相关性；冬季,三种嘴长的鸻鹬类与各类型底栖动物均未表现出相关性。春季,视觉连续型觅食策略鸻鹬类密度与甲壳类密度显著正相关,触觉连续型觅食策略鸻鹬类密度双壳类密度及底栖动物总密度显著正相关,而奔——停型觅食策略鸻鹬类密度与栖息地内的各类底栖动动物均无显著相关性；冬季,3种觅食策略鸻鹬类与各类底栖动物均未表现出明显的相关性。春、秋季各类别鸻鹬类比例与面上型和面下型底栖动物的比例相关性均不显著。觅食策略相同的物种或主要食物的组成上存在差异,或主食食物的种类较多。取食行为与食物的可得性关系密切,对于同一类别的食物它们更倾向于取食密度较高的种类。
Shorebirds (Charadriiformes) are typical waterbirds wading in wetlands, as a very important component of the coastal and lake ecological system, the number of shorebirds is a measure of wetlands value. The food provided by stopover sites along the flyway are vital for shorebirds to refuel and finish their migration to the breeding ground. Yancheng Nature Reserve is an important stopsite for shorebirds in the East Asian-Australasian Flyway (EAAF). In order to study the role of Yancheng Nature Reserve on supporting the migration of shorebirds and provide scientific data to shorebirds conservation. In Sheyang estuary, the macrobenthos are the main food resources of shorebirds, the paper studied the resources status of macrobenthos, analysed the niche of macrobenthos on the tidal flat and the effect of Spartina alterniflora invasion on them.in addition, studied the resources status of shorebirds during the northward and southward migration and wintering period and the relationship between macrobenthos and shorebirds in spring and winter. The main results were as follows:
1.We found a total of 16 species of macrobenthic on intertidal flats of Sheyang estuary, Yancheng Nature Reserve. The species numbers found at 4 sampling plots (intertidal zone with Spartina alterniflora, high, middle and low tidal flat without plants) were different. The habitat variation was much more important reason of macrobenthic community distribution than the season. The Potamocorbula ustulata and Macoma praetextadensity had broader niche breadths, however the Glaucomya chinensis and Mactra veneriformis were narrower. There were great differences in the niche overlaps among all the species. The niche breadth and niche overlaps had close relationship with the species spatio-temporal distribution and quantity.
2. We found a total of 22 species of macrobenthic on tidal flats in Sheyang estuary, Yancheng Nature Reserve. The compositions of species between 2 contrast team(area with Spartina alterniflora and area without Spartina alterniflora) of each group were different. The effect of Spartina alterniflora from different habitats was different: either reduced the number of species, density, biomass and biodiversity in intertidal flat or enhanced these indices except for biomass in creek, however, the effect of Spartina alterniflora invasion on all indices was inconsistent in supralittoral zone.
3. A total of 22 species were observed at Sheyang Estuary, Yancheng Nature Reserve., we can found all spcies in spring,13 species can be found in autumn, there were only 8 species in winter. The Dunlin(Calidris alpina) was the dominant species in 3 seasons and the percentage of number of shorebirds was 3:1:1.5 in spring, autumn and winter. The density of shorebirds existed significant difference among habitats in spring, autumn and winter; the great number of species and Shannon-weinner diversity index were all found at creek in 3 seasons, The density of shorebirds was the much bigger at the high tidal flat without plants, middle tidal flat without plants and creek, the number of shorebirds were the biggest at high and middle tidal flat without plants; the shorebirds were also found at aquaculture ponds, but the density was very low and most of them inhabited in there. The migratory peak period was the last ten to twenty day in April, and finish at the last ten to twenty day in May, different species had few difference at the migratory peak period.
4. The habitat with higher density of macrobenthic had much more shorebirds. the densities of shorebirds with bill-length beyond 50mm were remarkable positive correlation with the densities of Crustacean, the densities of shorebirds with bill-length below 25mm were remarkable positive correlation with the densities of Bivalve, however, the densities of shorebirds with bill-length between 25-50mm weren't remarkable correlation with the densities of several forms of macrobenthic in spring; but the densities of shorebirds with 3 kind of bill-length weren't remarkable correlation with the densities of several forms of macrobenthic in winter. The densities of visual continuous shorebirds were remarkable positive correlated with the densities of Crustacean, the densities of Tactile continuous shorebirds were remarkable positive correlation with the densities of Bivalve, however, the densities of Pause-travel shorebirds weren't remarkable correlation with the densities of several forms of macrobenthic in spring; 3 foraging strategy shorebirds weren't remarkable correlation with the densities of every forms of macrobenthic in winter. The percentages of each categories of shorebirds weren't remarkablely correlated to the percentages of epifauna and infauna life forms. The species with same foraging strategy had either different main food or many kinds of food. the gained food affect foraging behaviour of shorebirds, for same type food, they are apt to select the species with high density.
1、研究地潮间带共有大型底栖动物16种,各生境的物种数不同。生境条件与潮间带大型底栖动物群落组成结构有一定的关系,同一生境大型底栖动物群落季节间差异不显著。焦河蓝蛤(Potamocorbula ustulata)和明细白樱蛤(Macoma praetexta)的生态位较宽,中国绿螂(Glaucomya chinensis)和四角蛤蜊(Mactra veneriformis)较窄。不同的物种间生态位重叠值差异较大。物种生态位宽度、物种之间的生态位重叠值与物种的时空分布和数量密切相关。
2、研究地滩涂共发现大型底栖动物22种。米草区和非米草区的物种组成不同,不同生境中的互花米草对底栖动物的影响不甚相同：潮间带互花米草的入侵降低了大型底栖动物的物种数、密度、生物量和多样性；潮沟米草的入侵提高了大型底栖动物的物种数、密度和多样性；而潮上带米草入侵对大型底栖动物的各指标影响不一。
3、研究地内共统计到鸻鹬类22种,这些种类在春季均能看到,秋季13种,冬季仅8种；黑腹滨鹬(Calidris alpina)是3个季节共同的优势种,春季、秋季和冬鸻鹬类的数量比值约为3：1：1.5；3个季节鸻鹬类的密度在生境间的差异均显著；3个季节,鸻鹬类物种数和Shannon-weinner多样性指数最多的均是潮沟,密度较高的是高潮带光滩、中潮带光滩和潮沟,数量最多的高潮带光滩和中潮带光滩；养殖塘也有一定数量的鸻鹬类分布,但密度较低,多以栖息为主；春季多数鸟类的迁徙高峰在4月中下旬,结束于5月中下旬,不同的类群迁徙高峰略有差异。
4、底栖动物密度较高的生境,鸻鹬类的密度也较高。春季,嘴长50mmm以上鸻鹬类的密度与甲壳类密度显著正相关,嘴长25mm以下鸻鹬类密度与双壳类密度及底栖动物总密度显著正相关,嘴长25-50mm鸻鹬类密度与任何一类底栖动物均未表现出明显相关性；冬季,三种嘴长的鸻鹬类与各类型底栖动物均未表现出相关性。春季,视觉连续型觅食策略鸻鹬类密度与甲壳类密度显著正相关,触觉连续型觅食策略鸻鹬类密度双壳类密度及底栖动物总密度显著正相关,而奔——停型觅食策略鸻鹬类密度与栖息地内的各类底栖动动物均无显著相关性；冬季,3种觅食策略鸻鹬类与各类底栖动物均未表现出明显的相关性。春、秋季各类别鸻鹬类比例与面上型和面下型底栖动物的比例相关性均不显著。觅食策略相同的物种或主要食物的组成上存在差异,或主食食物的种类较多。取食行为与食物的可得性关系密切,对于同一类别的食物它们更倾向于取食密度较高的种类。
Shorebirds (Charadriiformes) are typical waterbirds wading in wetlands, as a very important component of the coastal and lake ecological system, the number of shorebirds is a measure of wetlands value. The food provided by stopover sites along the flyway are vital for shorebirds to refuel and finish their migration to the breeding ground. Yancheng Nature Reserve is an important stopsite for shorebirds in the East Asian-Australasian Flyway (EAAF). In order to study the role of Yancheng Nature Reserve on supporting the migration of shorebirds and provide scientific data to shorebirds conservation. In Sheyang estuary, the macrobenthos are the main food resources of shorebirds, the paper studied the resources status of macrobenthos, analysed the niche of macrobenthos on the tidal flat and the effect of Spartina alterniflora invasion on them.in addition, studied the resources status of shorebirds during the northward and southward migration and wintering period and the relationship between macrobenthos and shorebirds in spring and winter. The main results were as follows:
1.We found a total of 16 species of macrobenthic on intertidal flats of Sheyang estuary, Yancheng Nature Reserve. The species numbers found at 4 sampling plots (intertidal zone with Spartina alterniflora, high, middle and low tidal flat without plants) were different. The habitat variation was much more important reason of macrobenthic community distribution than the season. The Potamocorbula ustulata and Macoma praetextadensity had broader niche breadths, however the Glaucomya chinensis and Mactra veneriformis were narrower. There were great differences in the niche overlaps among all the species. The niche breadth and niche overlaps had close relationship with the species spatio-temporal distribution and quantity.
2. We found a total of 22 species of macrobenthic on tidal flats in Sheyang estuary, Yancheng Nature Reserve. The compositions of species between 2 contrast team(area with Spartina alterniflora and area without Spartina alterniflora) of each group were different. The effect of Spartina alterniflora from different habitats was different: either reduced the number of species, density, biomass and biodiversity in intertidal flat or enhanced these indices except for biomass in creek, however, the effect of Spartina alterniflora invasion on all indices was inconsistent in supralittoral zone.
3. A total of 22 species were observed at Sheyang Estuary, Yancheng Nature Reserve., we can found all spcies in spring,13 species can be found in autumn, there were only 8 species in winter. The Dunlin(Calidris alpina) was the dominant species in 3 seasons and the percentage of number of shorebirds was 3:1:1.5 in spring, autumn and winter. The density of shorebirds existed significant difference among habitats in spring, autumn and winter; the great number of species and Shannon-weinner diversity index were all found at creek in 3 seasons, The density of shorebirds was the much bigger at the high tidal flat without plants, middle tidal flat without plants and creek, the number of shorebirds were the biggest at high and middle tidal flat without plants; the shorebirds were also found at aquaculture ponds, but the density was very low and most of them inhabited in there. The migratory peak period was the last ten to twenty day in April, and finish at the last ten to twenty day in May, different species had few difference at the migratory peak period.
4. The habitat with higher density of macrobenthic had much more shorebirds. the densities of shorebirds with bill-length beyond 50mm were remarkable positive correlation with the densities of Crustacean, the densities of shorebirds with bill-length below 25mm were remarkable positive correlation with the densities of Bivalve, however, the densities of shorebirds with bill-length between 25-50mm weren't remarkable correlation with the densities of several forms of macrobenthic in spring; but the densities of shorebirds with 3 kind of bill-length weren't remarkable correlation with the densities of several forms of macrobenthic in winter. The densities of visual continuous shorebirds were remarkable positive correlated with the densities of Crustacean, the densities of Tactile continuous shorebirds were remarkable positive correlation with the densities of Bivalve, however, the densities of Pause-travel shorebirds weren't remarkable correlation with the densities of several forms of macrobenthic in spring; 3 foraging strategy shorebirds weren't remarkable correlation with the densities of every forms of macrobenthic in winter. The percentages of each categories of shorebirds weren't remarkablely correlated to the percentages of epifauna and infauna life forms. The species with same foraging strategy had either different main food or many kinds of food. the gained food affect foraging behaviour of shorebirds, for same type food, they are apt to select the species with high density.
引文
Ayers DR, Strong DR,2002. The Spartina invasion of San Francisco Bay[J].Aquatic Nuisance Species Digest, 4:37-39.
Burger J, Gochfeld M.1991. Human activity influence and diurnal and nocturnal foraging of sanderlings (Calldris alba) [J].The Condor,93:259-265.
Colwell MA, Taft OW.2000. Waterbird communities in managed wetlands of varying water depth. Waterbirds,23:45-55.
Dias MP, Dias JP, Granadeiro ML, et al.2006. Distance to high-tide roosts constrains the use of foraging areas by dunlins:Implications for the management of estuarine wetlands[J]. Biological conservation,131: 446-452
Duffy DC, Atkins N, Schneider DC.1981. Do shorebirds compete on their wintering grounds[J].Auk,98: 215-248.
Eybert MC, Thomas G, Sophie Q, et al.2003. Shorebird community variations indicative of a general perturbationin the Mont-Saint-Michelbay(France)[J]. C. R.Biologies,326:S140-S147
Nuka T, Norman CP, Kuwabara K, et al.2005.Feeding behavior and effect of prey availability on Sanderling Calidris alba distribution on Kujukuri Beach[J].Ornithol. Sci.,4,139-146.
Finn PG, Carla PC, Peter VD.2008.Prey versus substrate as determinants of habitat choice in a feeding shorebird[J]. Estuarine, Coastal and Shelf Science,1-10
Finn PG, Catterall CP, Driscoll P.2007. Determinants of preferred intertidal feeding habitat for Eastern Curlew: a study at two spatial scales[J].Austral Ecology,32,131e144.
Fujii. T.2007. Spatial patterns of benthic macrofauna in relation to environmental variables in an intertidal habitat in the Humber estuary, UK:Developing a tool for estuarine shoreline management[J]. Estuarine, Coastal and Shelf Science,75:101-119.
Gochfeld M, Burger J, Jehl JJR.1984.The classification of the shorebirds of the world. Burger J and Olla B L(eds). Shorebirds:Breeding behavior and population[M]. New York:Plenum Press,5:1-16.
Goss-Custard JD.1984. Intake rates and food supply in migrating and wintering shorebirds [A]. In:Burger J, Olla BL. Shorebirds:Mi-gration and Foraging Behavior [M]. New York:Plenum Press.233-270.
Grant J.1984. Sediment microtopography and shorebird foraging[J]. Marine Ecology Progress Series,19:293-296.
Halliwell DR, Bucher C, Fehr A, et al.2002. Protecting Aquatic Quality of Tuktut Nogait National Park & Lower Hornaday River,NWT.Prepared for Meteorological Service of Canada[A],Enviroment canada Yellowkinfe,NT,64pp
Higgins, Thiel H.1988. Introduction to the study of meiofauna[M]. Washington:Smithsonian Institution Press.
Honkoop PJC, Pearson GB, Lavaleye MSS, et al.2006. Spatial variation of the intertidal sediments and macrozoo-benthic assemblages along Eighty-mile Beach, Northwestern Australia[J]. Journal of Sea Research,55:278-291.
Isola CR, Colwell, MA, Taft, OW.2000. Interspecific differences in habitat use of shorebirds and waterfowl foraging in managed wetlands of California's San Joaquin Valley [J]. Waterbirds,23:196-203.
Jing K, MaZJ, Li B, et al.2007.Foraging strategies involved in habitat use of shorebirds at the intertidal area of Chongming Dongtan,China[J].Ecologiccal Research,22:559-570.
Kalejta B, Hockey PAR.1994. Distribution of Shorebirds at the Berg River estuary, South Africa, in relation to foraging mode, food supply and environmental features[J]. Ibis,136,233-239.
Kundu S, Mondal N, Lyla PS, et al.2010.Biodiversity and seasonal variation of macrobenthic infaunal community in the inshore waters of Parangipettai Coast[J]. Environment Monit Assess,163:67-79.
Lampadariou N, Akoumianaki I, Karakassis I.2008.Use of the size fractionation of the macrobenthic biomass for the rapid assessment of benthic organic enrichment[J].Ecological Indicators,8:729-742.
Li XZ, Wang JB, Wang HF, et al.2005a. Secondary Production of Macrobenthosfrom the East China Sea[J]. Chinese Journal of Applied and Environmental Biology,11 (4):459-462.
Li XZ, Zhang BL, Wang HF.2005b. Study on the secondary production of macrobenthos from the anchovy spawning ground in the Southern Yellow Sea[J]. Chinese Journal of Applied and Environmental Biology, 11(3):324-327.
Ma ZJ, Li B, Zhao B, et al.2004. Are artificial wetlands good alternatives to natural wetlandsforwaterbirds? —A case study on Chongming Island, China[J]. Biodiversity and Conservation,13:333-350.
Marzano CN, Liaci LS, Fianchini A, et al.2003.Distribution, persistence and change in the macrobenthos of the lagoon of Lesina(Apulia, southern Adriatic Sea)[J]. Ocesnoligica Acta,26:57-66.
McLachlan GR, Wooldridge T, Schramn M, et al.1980. Seasonal abundance, biomass and feeding of shorebirds on sandy beaches in the Eastern Cape, South Africa[J].Ostrich,51:44-52.
Mcnicol DK, Bendell BE, Ross RK.1987.Studies of the Effects of Acidification on Aquatic Wildlife in Canada:Waterfowl and Trophic Relationships in Small Lakes in Northernnn Ontario [A].Canadian Wildlife Service Occasional,62.
Metcalfe NB.1985. Prey detection by intertidally feeding Lapwing Zeits[J].Tierpsychologie,64:45-57.
Moreira F.1999.On the use by birds of intertidal areas of the Tagus estuary:implications for managemen[J]. Acquatic Ecology,33:301-309
Mucha AP, Costa MH.1999. Macrozoobenthic community structure in two Portuguese estuaries:Relationship with organic enrichment and nutrient gradients[J]. Acta Oecol,20:363-376.
Nebel S, Porter JL, Kinngsford RT.2008. Long-term trends of shorebird populations in eastern Australia and impacts of freshwater extraction[J].Biological Conservation,971-980
Ohtaka A, Watanabe R, Im S, et al.2010.Spatial and seasonal changes of net plankton and zoobenthos in Lake Tonle Sap,Cambodia[J]. Limnology,11:85-94.
Owner D, Rohweder DA.2003. Distribution and habitat of Pied Oyster-catchers (Haematopus longirostris) inhabiting ocean beaches in northern New South Wales[J]. Emu,103,163e169.
Peter RT, Wookdridge TH.2003. What limits the distribution of subtidal macrobenthos in permanently open and temporarily openΠclosed South African estuaries? Salinity vs. sediment paricle size. Estuarine[J]. Coastal and Shelf Science,57:225-238.
Prater AJ.1981. Estuary Birds of Britain and Ireland[M]. Calton:Poyser.
Puttick GM.1979. Foraging behavior and activity budgets of Curlew Sandpipers [J].Ardea,67:111-122.
Puttick GM.1984. Foraging and activity patterns in wintering shorebirds. In:Burger, J., Olla, B.L. (Eds.), Behavior of Marine Animals. Shorebirds:Migration and Foraging Behavior[M]. Plenum Press, New York and London,6:203-231.
Rogers DI, Battley PF, Theunis P, et al.2006b. High-tide habitat choice:insights from modelling roost selection by shorebirds around a tropical bay[J]. Animal Behaviour,72:563-575
Rogers DI, Theunis P, Chris JH.2006a. Roost availability may constrain shorebird distribution:Exploring the energetic costs of roosting and disturbance around a tropical bay[J]. Biological conservation,133:225-235
Rohweder DA, Baverstock PR.1996. Preliminary investigation of nocturnal habitat use by migratory waders (Order Charadriformes) in Northern New South Wales[J]. Wildlife Research,23:169-184.
Sanchez MI, Green AJ, Castellanos EM.2006. Spatial and temporal fluctuations in presence and use of chironomid prey by shorebirds in the Odiel saltpans,south-west Spain[J]. Hydrobiologia,567:329-340.
Sosuke O, Yasunori K, Kengo K, et al.2008.Relationship between macrobenthos and physical habitat characters in tidal flat in eastern Seto Inland Sea, Japan[J]. Marine Pollution Bulletin,57:142-148
Spruzen F L, Richardson A M M, Woehler E J.2008.Influence of environmental and prey variables on low tide shorebird habitat use within the Robbins Passage wetlands, Northwest Tasmania[J]. Estuarine, Coastal and Shelf Science,78:122-134
Stillman RA, Goss-Custard JD.2002. Seasonal changes in the response of oyster catchers Haematopus ostralegusto human disturbance [J] Journal of Avian Biology,33:358-365.
Straw P.1997.Shorebird conservation in the Aisa-Pacific region.Australasian Wader Study Group of Birds Australia,Melbourne,Australia
Summers R W, UnderhiilL G, Simpson A.2002. Habitat preference of waders Charadrii on the coast of Orkney island[J]. Bird Study,49:60-66.
Takekawa JY, Warnock N, Martinelli GM, et al.2002. Wterbirds use of Bayland in the San Francisco Bay estuary:Movements of long-billed Dowitchers during the winter[J]. Waterbirds,25(2):93-105.
Thompson JJ.1998. Interseasonal changes in shorebird habitat specialisation in Moreton Bay, Australia[J]. Emu,98:117-126.
Thrush SF, Hewitt JE, Gibbs M, et al.2006. Functional role of large organisms in intertidal communities: community effects and ecosystem function[J]. Ecosystem,9:1029-1040.
Webster PJ, Rowden AA, Attrill MJ.1998. Effect of shoot density on the infaunal macro-invertberate communitywith in a Zostera marina seagrass bed[J]. Estuarine, Coastland Shelf Science,47:351-357.
Wetlands International.2006.Waterbirds population estimates, fourthed. Wetlands international, Wageningen, The Netherlannds.
Zou FS, Zhang HH, Dahmer T, et al.2008.The effects of benthos and wetland area on shorebird abundance and species richness in coastal mangrove wetlands of Leizhou Peninsula, China [J]. Forest Ecology and Management,255:3813-3818.
安传光,赵云龙,林凌,等.2008.崇明岛潮间带夏季大型底栖动物多样性[J].生态学报,28(2)：577-586.
鲍毅新,葛宝明,郑祥,等.2007.温州湾灵昆岛东滩潮间带大型底栖动物群落的季节动态[J].水生生物学报,31(3)：437-444.
鲍毅新,胡知渊,李欢欢,等.2008.灵昆东滩围垦区内外大型底栖动物季节变动和功能群的比较[J].动物学报,53(3)：416-427.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M],北京：中国林业出版社.
陈中义,付萃长,王海毅,等.2005.互花米草入侵东滩盐沼对大型底栖无脊椎动物群落的影响[J].湿地科学,3(1)：1-7.
崔志兴,陈龙小.1998.长江口迁徙鸻形目鸟类群落结构十三年来的变化.见：社团法人台北市野鸟学会编.第三届海峡两岸鸟类学术研讨会论文集[A].台湾鸟类资迅社,189-200
葛宝明,鲍毅新,郑祥,等.2005.灵昆岛潮间带大型底栖动物群落结构与生态位分析[J].生态学报,25(11)：3037-3043.
葛宝明,鲍毅新,程宏毅,等.2008.灵昆岛东滩潮间带大型底栖动物功能群及营养等级构成[J].生态学报,28(10)：4796-4804.
葛振鸣,王天厚,施文或,等.2006a.长江口杭州湾鸻形目鸟类群落季节变化和生境选择[J].生态学报,26(1)：40-47
葛振鸣,王天厚,周晓,等.2006b.上海崇明东滩堤内次生人工湿地鸟类冬春季生境选择的因子分析[J].动物学研究,27(2)：144-150
葛振鸣,周晓,施文或,等.2007.九段沙湿地鸻形目鸟类迁徙季节环境容纳量[J].生态学报,27(1)：90-96
胡知渊,鲍毅新,葛宝明,等.2006.围垦滩涂潮沟秋季大型底栖动物群落和生态位分析[J].动物学报,52(4)：800-809.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报, 28(4)：1498-1507.
江红星,徐文彬,钱法文,等.2007.栖息地演变与人为干扰对升金湖越冬水鸟的影响[J].应用生态学报,18(8)：1832-1836
李宝泉,李新正,王洪法,等.2007.长江口附近海域大型底栖动物群落特征[J].动物学报,53(1)：76-82.
李冠国,范振刚.2004.海洋生态学[M].北京：高等教育出版社.
李新正,王洪法,张宝琳.2005.胶洲湾大型底栖动物次级生产力初探[J].海洋与湖沼,36(6)：527-533.
李新正,于海燕,王永强,等.2001.胶州湾大型底栖动物的物种多样性现状[J].生物多样性,9(1)：80-84.
廖一波,曾江宁,陈全震,等.2007.嵊泅海岛不同底质潮间带春秋季大型底栖动物的群落格局[J].动物学报,53(6)：1000-1010.
吕小梅,方少华,张跃平,等.2008.福建海坛海峡潮间带大型底栖动物群落结构及次级生产力[J].动物学报,54(3)：428-435.
全为民,赵云龙,朱江兴,等.2008.上海市潮滩湿地大型底栖动物的空间分布格局[J].生态学报,28(10)：5179-5187.
唐承佳,陆健健.2002.围垦堤内迁徙鸻鹬群落的生态学特性[J].动物学杂志,37(2)：27-33.
唐仕华,虞快.1998.长江口南岸春季鸻形目鸟类的迁徙研究[J].华东师范大学学报(自然科学版),12(动物学专辑)：135-139.
王天厚,钱国桢.1987.长江口及杭州湾北部滩涂的生物群落特征分析[J].生态学杂志,6(2)：35-38.
徐晓军,由文辉,张锦平,等.2008.崇明东滩底栖动物群落与潮滩高程的关系[J].江苏环境科技,21(3)：30-35.
杨洪燕,张正旺.2006.渤海湾地区红腹滨鹬迁徙动态的初步研究[J].动物学杂志,41(3)：85-89.
杨月伟,张伟,王军,等.2007.人为干扰对两种鸻形目鸟类觅食行为的影响[J].曲阜师范大学学报(自然科学版),33(3)：509-511.
杨月伟,夏贵荣,丁平,等.2005.人为干扰对黑腹滨鹬觅食行为的影响[J].动物学研究,26(2)：136-141
袁兴中.2001.河口潮滩湿地底栖动物群落的生态学研究[A].华东师范大学博士学位论文,上海
袁兴中,陆健健,刘红.2002.长江口底栖动物功能群分布格局及其变化[J].生态学报,22(12)：2054-2062.
赵锦霞,刘昊,张利权.2008.崇明东滩越冬鸟类在养殖塘的空间分布[J].动物学研究,29(2)：212-218.
赵延茂,吕卷章,朱书玉,等.2001.黄河三角洲自然保护区形目鸟类研究[J]动物学报,(47)：157-161.
郑荣泉,张永普,李灿阳,等.2007.乐清湾滩涂大型底栖动物群落结构的时空变化[J].动物学报,53(2)：390-398.
仲阳康,周慧,施文,等.2006.上海滩涂春季鸻形目鸟类群落及围垦后生境选择[J].长江流域资源与环境,15(3)：378-383.
周慧,仲阳康,赵平,等.2005.崇明东滩冬季水鸟生态位分析[J].动物学杂志,40(1)：59-65.
周进,李新正,李宝泉.2008.黄海中华哲水蚤度夏区大型底栖动物的次级生产力[J].动物学报,54(3)：436-441
周时强,郭丰,吴荔生,等.2001.福建海岛潮间带底栖生物群落生态的研究[J].海洋学报,23(5)：104-109.
邹发生,杨琼芳,蔡俊钦,等.2008.雷州半岛红树林湿地鸟类多样性[J].生态学杂志,27(3)：383-390
Ayers DR, Strong DR.2002.The Spartina invasion of San Francisco Bay[J]. Aquatic Nuisance Species Digest,4:37-39.
Boehs GT, Absher TH, Cruz-Kaled AD.2004.Composition and distribution of benthic molluscs on intertidal flats of Paranagua Bay(Parana, Brazil) [J]. Scientia Marina,68:537-543.
Bonsdorff E.2006. Zoobenthic diversity-gradients in the Baltic Sea:Continuous post-glacial succession in a stressed ecosystem[J]. Journal of Experimental Marine Biology and Ecology,330:383-391.
Edgar GT, Barrett NS.2002. Benthic macrofauna in Tasmanian estuaries:scales of distribution and relationships with environmental variables[J]. Journal of Experimental Marine Biology and Ecology, 270:1-24.
Feder HM, Jewett SC.2007. Blanchard AL.Southeastern Chukchi Sea (Alaska) macrobenthos[J]. Ploar Biology,30:261-275.
Fujii T.2007. Spatial patterns of benthic macrofauna in relation to environmental variables in an intertidal habitat in the Humber estuary, UK:Developing a tool for estuarine shoreline managemen[J]. Estuarine, Coastal and Shelf Science,75:101-119.
Honkoop PJC, Pearson GB, Lavaleye MSS, et al.2006. Spatial variation of the intertidal sediments and macrozoo-benthic assemblages along Eighty-mile Beach, Northwestern Australia[J]. Journal of Sea Research,55:278-291.
Peter RT, Wookdridge TH.2003. What limits the distribution of subtidal macrobenthos in permanently open and temporarily open/closed South African estuaries? Salinity vs. sediment paricle size[J]. Estuarine,Coastal and Shelf Science,57:225-238.
Spruzen FL, Richardson AMM, Woehler EJ.2008. Spatial variation of intertidal macroinvertebrates and environmental variables in Robbins Passage wetland, NW Tasmania[J]. Hydrobiologia,598:325-342.
Taft OW, Haig SM.2005. The value of agricultural wetlands as invertebrate resources for wintering shorebirds[J].Agriculture, Ecosystems and Environment,110:249-256.
Thrush SF, Hewitt JE, GibbsM, et al.2006.Functional role of large organisms in intertidal communities: community effects and ecosystem function[J]. Ecosystem,9:1029-1040.
Wieking G, Kroncke I.2005. Is benthic trophic structure affected by food quality? The Dogger Bank example[J]. Marine Biology,146:387-400.
鲍毅新,葛宝明,郑祥,等.2006.温州湾天河滩涂大型底栖动物群落分布与季节变化[J].动物学报,52(1)：45-52.
鲍毅新,胡知渊,李欢欢,等.2008.灵昆东滩围垦区内外大型底栖动物季节变化和功能群的比较[J].动物学报,54(3)：416-427.
葛宝明,鲍毅新,程宏毅,等.2008.灵昆岛东滩潮间带大型底栖动物功能群及营养等级构成[J].生态学报,28(10)：4796-4804.
葛宝明,鲍毅新,郑祥.2005.灵昆岛围垦滩涂潮沟大型底栖动物群落生态学研究[J].生态学报,25(3)：446-452.
葛宝明,郑祥,程宏毅,等.2007.灵昆岛围垦滩涂潮沟大型底栖动物群落和物种生态位分析[J].水生生物学报,31(5)：675-681.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报,28(4)：1498-1507.
金亮,蔡立哲,周细平,等.2007.深圳湾北岸泥滩大型底栖动物次级生产力研究[J].台湾海峡,26(3)：413-421.
李欢欢,鲍毅新,胡知渊,等.2007.杭州湾南岸大桥建设区域潮间带大型底栖动物功能群及营养等级的季节动态[J].动物学报,53(6)：1011-1023.
李新正,于海燕,王永强,等.2001.胶州湾大型底栖动物的物种多样性现状[J].生物多样性,9(1)：80-84.
厉红梅,蔡立哲,林丽珠,等.2001.深圳湾潮间带底栖动物群落结构的等级聚类与非度量多维标度测序[J].厦门大学学报,40(3)：35-740.
刘建军,徐艳华.1997.水磨河底栖动物群落结构的生态位分析[J].水生生物学报,21(2)：101-108.
刘奕琳.2006.盐城海滨湿地生态系统的研究[A].博士学位论文.南京林业大学.
马藏允,刘海,王惠卿,等.1997.底栖动物群落结构变化多元变量统计分析[J].中国环境科学,17(4)：297-300.
马克平,刘玉明.1994.生物群落多样性的测度方法I.a多样性的测定方法(下)[J].生物多样 性,2(4)：231-239.
马克平.1994.生物多样性的测度方法.钱迎倩,马克平主编.生物多样性研究的原理与方法[M].北京：中国科学技术出版社,141-165.
任美锷,许廷官,朱季文,等.1986.江苏省海岸带和海涂资源综合调查报告[M].北京：海洋出版社.
王加连,吕士成.2008.江苏省盐城滩涂野生动物资源调查研究[J].四川动物,27(4)：620-625.
徐治国,何岩,闫百兴,等.2007.三江平原典型沼泽湿地植物种群的生态位[J].应用生态学报,18(4)：783-787.
袁兴中,陆健健.2001.长江口岛屿湿地的底栖动物资源研究[J].自然资源学报,16(1)：35-41.
袁兴中,陆健健,刘红.2002.长江口底栖动物功能群分布格局及其变化[J].生态学报,22(12)：2054-2062.
赵永强,曾江宁,高爱根,等.2009.椒江口潮间带大型底栖动物的生态位[J].应用生态学报,20(5)1176-1183.
郑荣泉,张永普,李灿阳,等.2007.乐清湾滩涂大型底栖动物群落结构的时空变化[J].动物学报,53(2)：390-398.
周晓,葛振鸣,施文或,等.2006.长江口九段沙湿地大型底栖动物群落结构的季节变化规律[J].应用生态学报,17(11)：2079-2083.
周晓,葛振鸣,施文或,等.2007.长江口新生湿地大型底栖动物群落时空变化格局[J].生态学杂志,26(3)372-377.
朱晓佳,钦佩.2003.外来种互花米草及米草生态工程[J].海洋科学,27(12)：14-19.
Angradi TR, Hagan SM, Able KW.2001. Vegetation type and the intertidal macro-invertebrate fauna of a brackish marsh:Phragmites vs. Spartina[J]. Wetland,21(9):75-92.
Cordell JR, Simenstad CA, Feist B, et al.1998.Ecological effects of S partina alterniflora invasion of the Littoral flat community in Willapa Bay,Washington[A].Abstracts from the Eighth International Zebra Mussel and Other Nuisance Species Conference, Sacramento California.
Curtis CD, Donald RS.1996. Status, prediction and prevention of introduced cordgrass Spartina spp. Invasion in pacific estuaries, USA[J]. Biological Conservation,78:51-58.
Hedge P, Kriwoken LK.2000. Evidence for effects of Spartina anglica invasion on macrobenthic fauna in Little Swanport estuary, Tasmania[J]. Austral Ecology,25:150-159.
Lana P, Guiss C.1991. Influence of Spartina alterniflora on structure and temporal variability of macrobenthic associations intidal flat of Paranagua Bay,Brazil[J]. Marine Ecolog Progress Series,9(73):231-234.
Luiting VT, Cordell JR, Olson AM,et al.1997.Doex exotic Spartina alterniflora change benthic invertebrate assemblages. In:Pattern K (ed.), Proceedings of the Second International Spartina Conference[A]. Washington State University,Olympia,48-50.
Neira C, Grosholz ED, Levin LA, et al.2006. Mechanisms generating modification of benthos following tidal flat invasion by a Spartina hybrid[J]. Ecological applications,16(17):1391-1404.
Posey MH, Alphin TD, Meyer DL, et al.2003.Benthic communities of common reed Phragmites australis and marsh cordgrass S partina alterniflora marshes in Chesapeake Bay[J]. Marine Ecology-Progress Series, 261:51-61.
Posey MH, Alphin TD, Meyer DL, et al.2003. Benthic communities of common reed Phragmites australis and marsh cordgrass Spartina alterniflora marshes in Chesapeake Bay[J]. Marine Ecology- Progress Series, 261:51-61.
Talley TS, Levin LA.2001. Modification of sediment s and macrofauna by an invasive marsh plant[J]. Biological Invasions,3:51-68.
Zhang RS, Shen YM, Lu LY, et al.2004. Formation of Spartina alterniflora salt marshes on the coast of Jiangsu Province[J].China.Ecol,23(2):95-105.
Zhao YJ, Zeng JN, Chen QZ, et al.2009. Community pattern of macrozoobenthos in distinct habitats with different densities of Spartina alterniflora Loisel[J]. Journal of Natural Resources,24(4):631-638.
Zhou HX, Liu JE, Qin P.2009. Impacts of an alien species(Spartina alterniflora) on the macrobenthos community of Jiangsu coastal inter-tidal ecosystem[J]. Ecological Engineering,35:521-528.
陈忠义,付萃长,王海毅,等.2005.互花米草入侵东滩盐沼对大型底栖无脊椎动物群落的影响[J].湿地科学,3(1)：1-7.
葛宝明,鲍毅新,郑祥.2005.灵昆岛围垦滩涂潮沟大型底栖动物群落生态学研究[J].生态学报,25(3)：446-452.
洪荣标,吕小梅,陈岚,等.2005.九龙江口红树林湿地与米草湿地的底栖生物[J].台湾海峡,24(2)：189-194.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报,28(4)：1498-1507.
田家怡,申保忠,李建庆,等.2009.黄河三角洲外来入侵物种米草对滩涂底栖动物的影响[J].海洋环境科学,28(6)：687-690.
谢文静,高抒.2009.江苏中部海岸互花米草盐沼底栖动物的组成特征[J].南京大学学报(自然科学),45(4)：553-560.
谢志发,何文珊,刘文亮,等.2008.不同发育时间的互花米草盐沼对大型底栖动物群落的影响[J].生态学杂志,27(1)：63-67.
谢志发,章飞军,刘文亮,等.2007.长江口互花米草生长区大型底栖动物的群落特征(英文)[J].动物学研究.28(2)：167-171.
任美锷,许廷官,朱季文,等.1986.江苏省海岸带和海涂资源综合调查报告[M].北京：海洋出版社.
徐炳声.1999.上海植物志(下卷)[M]上海：上海科学技术文献出版社.
徐晓军,王华,由文辉,等.2006.崇明东滩互花米草群落中底栖动物群落动态的初步研究[J].海洋湖沼通报,2：89-95.
杨泽华,童春富,陆健健.2007.盐沼植物对大型底栖动物群落的影响[J].生态学报,27(11)：4387-4393.
虞蔚岩,李朝晖,华春,等.2009a.江苏盐城东台互花米草滩涂底栖无脊椎动物的多样性分析[J].海洋湖沼通报.1(6)：123-128.
虞蔚岩,李朝晖,黄成,等.2009b.互花米草入侵对盐城东台盐滩多毛纲动物多样性的影响及管理对策[J].上海海洋大学学报,18(1)：47-5.
赵永强,曾江宁,陈全震,等.2009.不同互花米草(Spartina alterniflora Loisel)密度生境中大型底栖动物群落格局[J].自然资源学报,24(4)：631-638.
仲崇信.1985.大米草简史及国外研究概况[A].南京大学学报(米草研究的进展-22年来的研究成果论文集),40(2)：1-30.
周晓,王天厚,葛振鸣,等.2006.长江口九段沙湿地不同生境中大型底栖动物群落结构特征分析[J].生物多样性,14(2)：165-171.
周晓,王天厚,葛振鸣,等.2006.长江口九段沙湿地不同生境中大型底栖动物群落结构特征分析[J].生物多样性,14(2)：165-171.
朱晓佳,钦佩.2003.外来种互花米草及米草生态工程[J].海洋科学,27(12)：14-19.
Battley PF, Piersma T.2005. Body composition and flight ranges of Bar-tailed Godwit (Limosa lapponica Baueri) from New Zealand[J]. Auk,122(3):922-937.
Battley PF, Piersma T, Rogers DI, et al.2004.Do body condition and plumage during fuelling predict northward departure dates of Great knots Calidris tenuirostris from northwest Australia? [J] Ibis,146:46-60.
Danufsky T, Colwell MA.2003.Winter shorebird communities and tidal flat characteristics at Humbodt Bay, Califonia[J]. The Condor,105:117-129.
Gan XJ, Cai YT, Choi CY, et al.2009. Potential impacts of invasive Spartina alterniflora on spring bird communities at Chongming Dongtan, a Chinese wetland of international importance [J].Estuarine,Coastal and Shelf Science,83:211-218.
Howes J, Bakewell D.1989.Shorebird Studies Manual. Kuala Lumpur:AWB Publication.143-147.
Ma ZJ, Li B, Zhao B, et al.2004. Are artificial wetlands good alternatives to natural wetlands for waterbirds? —A case study on Chongming Island, China[J]. Biodiversity and Conservation,13:333-350.
Piersma T, DeGoeij P, Tulp I.1993. An evalution of intertidal feeding habitats from a shorebird perspective towards relevant comparisons between temperate and tropical mudflats Netherlands [J] Journal of Sea Research,31:103-123.
Shepherd PCF, Lank DB.2004. Marine and agricultural habitat preferences of dunlin wintering in British Columbia[J]. Journal of Wildlife Management,68(1):61-73.
Weber LM, Haig SM.1997. Shorebird diet and size selection of nereid polychaetes in South Carolina coastal diked wetlands[J]. Journal of Field Omithology.68(3):358-366
Wilson JR, Barter M.1998. Identification of potentially important staging area for 'long jump' migrant waders in the east Asian-Australian flyway during northward migration[J]. Silt,32:16-26.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M].北京：中国林业出版社.
崔志兴,钱国祯,祝龙彪,等.1985.鸻形目鸟类的食性研究[J]动物学研究(增刊).6(4)：43-51.
葛振鸣,王天厚,施文或,等.2006a.长江口杭州湾鸻形目鸟类群落季节变化和生境选择[J].生态学报,26(1)：40-47.
葛振鸣,王天厚,周晓等.2006b.上海崇明东滩堤内次生人工湿地鸟类冬春季生境选择的因子分析[J].动物学研究,27(2)：144-150.
葛振鸣,周晓,施文或,等.2007.九段沙湿地鸻形目鸟类迁徙季节环境容纳量生态学报[J].27(1)：90-96.
国家环保总局.2003.优先保护17个生物多样性关键地区[J].生态经济,2：78.
金杰锋,刘伯锋,余希,等.2008.福建省兴化湾滨海养殖塘冬季水鸟的栖息地利用[J].动物学杂志,43(6)：17-24.
敬凯.2005.上海崇明东滩鸻鹬类中途停歇生态学研究.复旦大学(博士论文).
李鑫,吴建平,朱成伟,等.2010.卧龙湖自然保护区水鸟迁徙研究[J].野生动物,31(3)：135-138.
刘伯锋.2003.福建沿海湿地鸻鹬类资源调查[J].动物学杂志,38(6)：72-75.
马敬能,菲利普斯,何芬奇.2000.中国鸟类野外手册[J].湖南教育出版社.
唐承佳,陆健健.2002.围垦堤内迁徙鸻鹬群落的生态学特性[J].动物学杂志,37(2)：27-33.
王会,杜进进.1993.射阳盐场湿地禽类资源考察初报[J].动物学杂志,28(4)：21-24.
张国钢,梁伟,刘冬平,等.2005.海南岛越冬水鸟资源状况调查[J].动物学杂志,40(2)：80-85.
朱晶,敬凯,干晓静,等.2007.迁徙停歇期鸻鹬类在崇明东滩潮间带的食物分布[J].生态学报.27(6)：2149-2159.
Barbosa A, Moreno E.1999.Evolution of foraging strategies in shorebirds:an ecomophorlogical approach[J]. Auk,116:712-725.
Boettcher R, Haig SM, Bridges WC.1995. Habitat-related factors affecting the distribution of nonbreeding Ameriean Avocets in Coastal South-Carolina[J]. Condor,97:68-81.
Bryant, DM.1979. Effects of prey density and site character on estuary usage by overwintering waders (Charadrii) [J]. Estuarine and Coastal Marine Science,9:369-384.
Butler RW, Davidson NC, Morrison RIG.2001.Global-scale shorebird distribution in relation to productivity of near-shore ocean waters[J]. Waterbirds,24:224-232.
Davis CA, Smith LM.2001. Foraging strategies and niche dynamics of coexisting shorebirds at stopover sites in the southern Great Plains[J] Auk,118:484-495.
Davis CA,Smith LM.2001.Foraging strategies and niche dynamics of coexisting shorebirds at stopover sites in the southern Great Plains[J].Auk,118:484-495.
Goss-Custard JD.1977. The ecology of the Wash. Ⅲ. Density-related behaviour and the possible effects of a loss of feeding grounds on wading birds (Charadrii) [J]. Journal of Applied Ecology,14:721-739.
Jing K, Ma ZJ, Li B,et al.2007.Foraging strategies involved in habitat use of shorebirds at the intertidal area of Chongming Dongtan,China[J].Ecologiccal Research,22:559-570.
Kalejta B, Hockey PAR.1994. Distribution of Shorebirds at the Berg River estuary, South Africa, in relation to foraging mode, food supply and environmental features[J]. Ibis,136:233-239.
Moreira F.1999. On the use by birds of intertidal areas of the Tagus estuary:implications for management[J]. Aquatic Ecology 33:301-309.
Morrison RIG.1984 Migration system of some new world shorebirds.In:Olla BL(ed)Shorebirds:migration and foraging behavior[M].Plenum Press:New York,125-202.
Nuka T, Norman CP, Kuwabara K, et al.2005.Feeding behavior and effect of prey availability on Sanderling Calidris alba distribution on Kujukuri Beach[J].Omithol. Sci,4:139-146.
Placyk JSJ, Harrington BA.2004 Prey abundance and habitat use by migratory shorebirds at coastal stopover sites in Connecticut[J]. Journal of Field Ornithology,75:223-231.
Ribeiro PD, Iribarne OO, Navarro D, et al.2004. Environmental heterogeneity, spatial segregation of prey, and the utilisation of southwest Atlantic mudflats by migratory shorebirds[J]. Ibis 146:672-683.
Ribeiro PD, Iribarne 00, Navarro D, et al.2004. Environmental heterogeneity, spatial segregation of prey, and the utilisation of southwest Atlantic mudflats by migratory shorebirds [J]. Ibis,146:672-683.
Rogers D I.2002. What determines shorebirds feeding distribution in Roebuck Bay?[C].M Pepping et al.(eds).Intertidal sediments and benthic animals of Roebuck Bay Western Australia. Texel:Nertherland Institute for Sea Research(NIOZ),145-174.
Sanchez MI, Green AJ, Castellanos EM.2006. Spatial and temporal fluctuations in presence and use of chironomid prey by shorebirds in the Odiel saltpans,south-west Spain[J]. Hydrobiologia,567:329-340.
Schoener TW.1965. The evolution of bill size differences among sympatric congeneric species of birds[J].Evolution,19:189-213.
Spruzen FL, Richardson AMM, Woehler EJ.2008. Influence of environmental and prey variables on low tide shorebird habitat use within the Robbins Passage wetlands, Northwest Tasmania[J]. Estuarine, Coastal and Shelf Science,78:122-134.
Zharikov Y, Skilleter GA.2002. Sex-specific intertidal habitat use in subtropically wintering Bar-tailed Godwits[J]. Canadian Journal of Zoology. (Revue Canadienne de Zoologie),80:1918-1929.
Zou FS, Zhang HH, Dahmer T, et al.2008.The effects of benthos and wetland area on shorebird abundance and species richness in coastal mangrove wetlands of Leizhou Peninsula, China [J]. Forest Ecology and Management,255:3813-3818.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M].北京：中国林业出版社.
黑龙江省野生动物研究所.1992.黑龙江省鸟类志[M].北京：中国林业出版社.136-165.
孙儒泳.2001.动物生态学原理[M].北京：北京师范大学出版社.
唐仕华,虞快.1996.崇明东滩鸻形目鸟类群落及其食性研究[J].华东师范大学学报(自然科学版)动物学专辑,79-83.
袁兴中,陆健健.2001.长江口潮沟大型底栖动物群落的初步研究[J].动物学研究,22：211-215.
朱书玉,吕卷章,赵长征,等.2000.黄河三角洲国家级自然保护区鸻形目鸟类食性及觅食地的研究[J].山东林业科技,130：10-13.
Burger J, Gochfeld M.1991. Human activity influence and diurnal and nocturnal foraging of sanderlings (Calldris alba) [J].The Condor,93:259-265.
Colwell MA, Taft OW.2000. Waterbird communities in managed wetlands of varying water depth. Waterbirds,23:45-55.
Dias MP, Dias JP, Granadeiro ML, et al.2006. Distance to high-tide roosts constrains the use of foraging areas by dunlins:Implications for the management of estuarine wetlands[J]. Biological conservation,131: 446-452
Duffy DC, Atkins N, Schneider DC.1981. Do shorebirds compete on their wintering grounds[J].Auk,98: 215-248.
Eybert MC, Thomas G, Sophie Q, et al.2003. Shorebird community variations indicative of a general perturbationin the Mont-Saint-Michelbay(France)[J]. C. R.Biologies,326:S140-S147
Nuka T, Norman CP, Kuwabara K, et al.2005.Feeding behavior and effect of prey availability on Sanderling Calidris alba distribution on Kujukuri Beach[J].Ornithol. Sci.,4,139-146.
Finn PG, Carla PC, Peter VD.2008.Prey versus substrate as determinants of habitat choice in a feeding shorebird[J]. Estuarine, Coastal and Shelf Science,1-10
Finn PG, Catterall CP, Driscoll P.2007. Determinants of preferred intertidal feeding habitat for Eastern Curlew: a study at two spatial scales[J].Austral Ecology,32,131e144.
Fujii. T.2007. Spatial patterns of benthic macrofauna in relation to environmental variables in an intertidal habitat in the Humber estuary, UK:Developing a tool for estuarine shoreline management[J]. Estuarine, Coastal and Shelf Science,75:101-119.
Gochfeld M, Burger J, Jehl JJR.1984.The classification of the shorebirds of the world. Burger J and Olla B L(eds). Shorebirds:Breeding behavior and population[M]. New York:Plenum Press,5:1-16.
Goss-Custard JD.1984. Intake rates and food supply in migrating and wintering shorebirds [A]. In:Burger J, Olla BL. Shorebirds:Mi-gration and Foraging Behavior [M]. New York:Plenum Press.233-270.
Grant J.1984. Sediment microtopography and shorebird foraging[J]. Marine Ecology Progress Series,19:293-296.
Halliwell DR, Bucher C, Fehr A, et al.2002. Protecting Aquatic Quality of Tuktut Nogait National Park & Lower Hornaday River,NWT.Prepared for Meteorological Service of Canada[A],Enviroment canada Yellowkinfe,NT,64pp
Higgins, Thiel H.1988. Introduction to the study of meiofauna[M]. Washington:Smithsonian Institution Press.
Honkoop PJC, Pearson GB, Lavaleye MSS, et al.2006. Spatial variation of the intertidal sediments and macrozoo-benthic assemblages along Eighty-mile Beach, Northwestern Australia[J]. Journal of Sea Research,55:278-291.
Isola CR, Colwell, MA, Taft, OW.2000. Interspecific differences in habitat use of shorebirds and waterfowl foraging in managed wetlands of California's San Joaquin Valley [J]. Waterbirds,23:196-203.
Jing K, MaZJ, Li B, et al.2007.Foraging strategies involved in habitat use of shorebirds at the intertidal area of Chongming Dongtan,China[J].Ecologiccal Research,22:559-570.
Kalejta B, Hockey PAR.1994. Distribution of Shorebirds at the Berg River estuary, South Africa, in relation to foraging mode, food supply and environmental features[J]. Ibis,136,233-239.
Kundu S, Mondal N, Lyla PS, et al.2010.Biodiversity and seasonal variation of macrobenthic infaunal community in the inshore waters of Parangipettai Coast[J]. Environment Monit Assess,163:67-79.
Lampadariou N, Akoumianaki I, Karakassis I.2008.Use of the size fractionation of the macrobenthic biomass for the rapid assessment of benthic organic enrichment[J].Ecological Indicators,8:729-742.
Li XZ, Wang JB, Wang HF, et al.2005a. Secondary Production of Macrobenthosfrom the East China Sea[J]. Chinese Journal of Applied and Environmental Biology,11 (4):459-462.
Li XZ, Zhang BL, Wang HF.2005b. Study on the secondary production of macrobenthos from the anchovy spawning ground in the Southern Yellow Sea[J]. Chinese Journal of Applied and Environmental Biology, 11(3):324-327.
Ma ZJ, Li B, Zhao B, et al.2004. Are artificial wetlands good alternatives to natural wetlandsforwaterbirds? —A case study on Chongming Island, China[J]. Biodiversity and Conservation,13:333-350.
Marzano CN, Liaci LS, Fianchini A, et al.2003.Distribution, persistence and change in the macrobenthos of the lagoon of Lesina(Apulia, southern Adriatic Sea)[J]. Ocesnoligica Acta,26:57-66.
McLachlan GR, Wooldridge T, Schramn M, et al.1980. Seasonal abundance, biomass and feeding of shorebirds on sandy beaches in the Eastern Cape, South Africa[J].Ostrich,51:44-52.
Mcnicol DK, Bendell BE, Ross RK.1987.Studies of the Effects of Acidification on Aquatic Wildlife in Canada:Waterfowl and Trophic Relationships in Small Lakes in Northernnn Ontario [A].Canadian Wildlife Service Occasional,62.
Metcalfe NB.1985. Prey detection by intertidally feeding Lapwing Zeits[J].Tierpsychologie,64:45-57.
Moreira F.1999.On the use by birds of intertidal areas of the Tagus estuary:implications for managemen[J]. Acquatic Ecology,33:301-309
Mucha AP, Costa MH.1999. Macrozoobenthic community structure in two Portuguese estuaries:Relationship with organic enrichment and nutrient gradients[J]. Acta Oecol,20:363-376.
Nebel S, Porter JL, Kinngsford RT.2008. Long-term trends of shorebird populations in eastern Australia and impacts of freshwater extraction[J].Biological Conservation,971-980
Ohtaka A, Watanabe R, Im S, et al.2010.Spatial and seasonal changes of net plankton and zoobenthos in Lake Tonle Sap,Cambodia[J]. Limnology,11:85-94.
Owner D, Rohweder DA.2003. Distribution and habitat of Pied Oyster-catchers (Haematopus longirostris) inhabiting ocean beaches in northern New South Wales[J]. Emu,103,163e169.
Peter RT, Wookdridge TH.2003. What limits the distribution of subtidal macrobenthos in permanently open and temporarily openΠclosed South African estuaries? Salinity vs. sediment paricle size. Estuarine[J]. Coastal and Shelf Science,57:225-238.
Prater AJ.1981. Estuary Birds of Britain and Ireland[M]. Calton:Poyser.
Puttick GM.1979. Foraging behavior and activity budgets of Curlew Sandpipers [J].Ardea,67:111-122.
Puttick GM.1984. Foraging and activity patterns in wintering shorebirds. In:Burger, J., Olla, B.L. (Eds.), Behavior of Marine Animals. Shorebirds:Migration and Foraging Behavior[M]. Plenum Press, New York and London,6:203-231.
Rogers DI, Battley PF, Theunis P, et al.2006b. High-tide habitat choice:insights from modelling roost selection by shorebirds around a tropical bay[J]. Animal Behaviour,72:563-575
Rogers DI, Theunis P, Chris JH.2006a. Roost availability may constrain shorebird distribution:Exploring the energetic costs of roosting and disturbance around a tropical bay[J]. Biological conservation,133:225-235
Rohweder DA, Baverstock PR.1996. Preliminary investigation of nocturnal habitat use by migratory waders (Order Charadriformes) in Northern New South Wales[J]. Wildlife Research,23:169-184.
Sanchez MI, Green AJ, Castellanos EM.2006. Spatial and temporal fluctuations in presence and use of chironomid prey by shorebirds in the Odiel saltpans,south-west Spain[J]. Hydrobiologia,567:329-340.
Sosuke O, Yasunori K, Kengo K, et al.2008.Relationship between macrobenthos and physical habitat characters in tidal flat in eastern Seto Inland Sea, Japan[J]. Marine Pollution Bulletin,57:142-148
Spruzen F L, Richardson A M M, Woehler E J.2008.Influence of environmental and prey variables on low tide shorebird habitat use within the Robbins Passage wetlands, Northwest Tasmania[J]. Estuarine, Coastal and Shelf Science,78:122-134
Stillman RA, Goss-Custard JD.2002. Seasonal changes in the response of oyster catchers Haematopus ostralegusto human disturbance [J] Journal of Avian Biology,33:358-365.
Straw P.1997.Shorebird conservation in the Aisa-Pacific region.Australasian Wader Study Group of Birds Australia,Melbourne,Australia
Summers R W, UnderhiilL G, Simpson A.2002. Habitat preference of waders Charadrii on the coast of Orkney island[J]. Bird Study,49:60-66.
Takekawa JY, Warnock N, Martinelli GM, et al.2002. Wterbirds use of Bayland in the San Francisco Bay estuary:Movements of long-billed Dowitchers during the winter[J]. Waterbirds,25(2):93-105.
Thompson JJ.1998. Interseasonal changes in shorebird habitat specialisation in Moreton Bay, Australia[J]. Emu,98:117-126.
Thrush SF, Hewitt JE, Gibbs M, et al.2006. Functional role of large organisms in intertidal communities: community effects and ecosystem function[J]. Ecosystem,9:1029-1040.
Webster PJ, Rowden AA, Attrill MJ.1998. Effect of shoot density on the infaunal macro-invertberate communitywith in a Zostera marina seagrass bed[J]. Estuarine, Coastland Shelf Science,47:351-357.
Wetlands International.2006.Waterbirds population estimates, fourthed. Wetlands international, Wageningen, The Netherlannds.
Zou FS, Zhang HH, Dahmer T, et al.2008.The effects of benthos and wetland area on shorebird abundance and species richness in coastal mangrove wetlands of Leizhou Peninsula, China [J]. Forest Ecology and Management,255:3813-3818.
安传光,赵云龙,林凌,等.2008.崇明岛潮间带夏季大型底栖动物多样性[J].生态学报,28(2)：577-586.
鲍毅新,葛宝明,郑祥,等.2007.温州湾灵昆岛东滩潮间带大型底栖动物群落的季节动态[J].水生生物学报,31(3)：437-444.
鲍毅新,胡知渊,李欢欢,等.2008.灵昆东滩围垦区内外大型底栖动物季节变动和功能群的比较[J].动物学报,53(3)：416-427.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M],北京：中国林业出版社.
陈中义,付萃长,王海毅,等.2005.互花米草入侵东滩盐沼对大型底栖无脊椎动物群落的影响[J].湿地科学,3(1)：1-7.
崔志兴,陈龙小.1998.长江口迁徙鸻形目鸟类群落结构十三年来的变化.见：社团法人台北市野鸟学会编.第三届海峡两岸鸟类学术研讨会论文集[A].台湾鸟类资迅社,189-200
葛宝明,鲍毅新,郑祥,等.2005.灵昆岛潮间带大型底栖动物群落结构与生态位分析[J].生态学报,25(11)：3037-3043.
葛宝明,鲍毅新,程宏毅,等.2008.灵昆岛东滩潮间带大型底栖动物功能群及营养等级构成[J].生态学报,28(10)：4796-4804.
葛振鸣,王天厚,施文或,等.2006a.长江口杭州湾鸻形目鸟类群落季节变化和生境选择[J].生态学报,26(1)：40-47
葛振鸣,王天厚,周晓,等.2006b.上海崇明东滩堤内次生人工湿地鸟类冬春季生境选择的因子分析[J].动物学研究,27(2)：144-150
葛振鸣,周晓,施文或,等.2007.九段沙湿地鸻形目鸟类迁徙季节环境容纳量[J].生态学报,27(1)：90-96
胡知渊,鲍毅新,葛宝明,等.2006.围垦滩涂潮沟秋季大型底栖动物群落和生态位分析[J].动物学报,52(4)：800-809.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报, 28(4)：1498-1507.
江红星,徐文彬,钱法文,等.2007.栖息地演变与人为干扰对升金湖越冬水鸟的影响[J].应用生态学报,18(8)：1832-1836
李宝泉,李新正,王洪法,等.2007.长江口附近海域大型底栖动物群落特征[J].动物学报,53(1)：76-82.
李冠国,范振刚.2004.海洋生态学[M].北京：高等教育出版社.
李新正,王洪法,张宝琳.2005.胶洲湾大型底栖动物次级生产力初探[J].海洋与湖沼,36(6)：527-533.
李新正,于海燕,王永强,等.2001.胶州湾大型底栖动物的物种多样性现状[J].生物多样性,9(1)：80-84.
廖一波,曾江宁,陈全震,等.2007.嵊泅海岛不同底质潮间带春秋季大型底栖动物的群落格局[J].动物学报,53(6)：1000-1010.
吕小梅,方少华,张跃平,等.2008.福建海坛海峡潮间带大型底栖动物群落结构及次级生产力[J].动物学报,54(3)：428-435.
全为民,赵云龙,朱江兴,等.2008.上海市潮滩湿地大型底栖动物的空间分布格局[J].生态学报,28(10)：5179-5187.
唐承佳,陆健健.2002.围垦堤内迁徙鸻鹬群落的生态学特性[J].动物学杂志,37(2)：27-33.
唐仕华,虞快.1998.长江口南岸春季鸻形目鸟类的迁徙研究[J].华东师范大学学报(自然科学版),12(动物学专辑)：135-139.
王天厚,钱国桢.1987.长江口及杭州湾北部滩涂的生物群落特征分析[J].生态学杂志,6(2)：35-38.
徐晓军,由文辉,张锦平,等.2008.崇明东滩底栖动物群落与潮滩高程的关系[J].江苏环境科技,21(3)：30-35.
杨洪燕,张正旺.2006.渤海湾地区红腹滨鹬迁徙动态的初步研究[J].动物学杂志,41(3)：85-89.
杨月伟,张伟,王军,等.2007.人为干扰对两种鸻形目鸟类觅食行为的影响[J].曲阜师范大学学报(自然科学版),33(3)：509-511.
杨月伟,夏贵荣,丁平,等.2005.人为干扰对黑腹滨鹬觅食行为的影响[J].动物学研究,26(2)：136-141
袁兴中.2001.河口潮滩湿地底栖动物群落的生态学研究[A].华东师范大学博士学位论文,上海
袁兴中,陆健健,刘红.2002.长江口底栖动物功能群分布格局及其变化[J].生态学报,22(12)：2054-2062.
赵锦霞,刘昊,张利权.2008.崇明东滩越冬鸟类在养殖塘的空间分布[J].动物学研究,29(2)：212-218.
赵延茂,吕卷章,朱书玉,等.2001.黄河三角洲自然保护区形目鸟类研究[J]动物学报,(47)：157-161.
郑荣泉,张永普,李灿阳,等.2007.乐清湾滩涂大型底栖动物群落结构的时空变化[J].动物学报,53(2)：390-398.
仲阳康,周慧,施文,等.2006.上海滩涂春季鸻形目鸟类群落及围垦后生境选择[J].长江流域资源与环境,15(3)：378-383.
周慧,仲阳康,赵平,等.2005.崇明东滩冬季水鸟生态位分析[J].动物学杂志,40(1)：59-65.
周进,李新正,李宝泉.2008.黄海中华哲水蚤度夏区大型底栖动物的次级生产力[J].动物学报,54(3)：436-441
周时强,郭丰,吴荔生,等.2001.福建海岛潮间带底栖生物群落生态的研究[J].海洋学报,23(5)：104-109.
邹发生,杨琼芳,蔡俊钦,等.2008.雷州半岛红树林湿地鸟类多样性[J].生态学杂志,27(3)：383-390
Ayers DR, Strong DR.2002.The Spartina invasion of San Francisco Bay[J]. Aquatic Nuisance Species Digest,4:37-39.
Boehs GT, Absher TH, Cruz-Kaled AD.2004.Composition and distribution of benthic molluscs on intertidal flats of Paranagua Bay(Parana, Brazil) [J]. Scientia Marina,68:537-543.
Bonsdorff E.2006. Zoobenthic diversity-gradients in the Baltic Sea:Continuous post-glacial succession in a stressed ecosystem[J]. Journal of Experimental Marine Biology and Ecology,330:383-391.
Edgar GT, Barrett NS.2002. Benthic macrofauna in Tasmanian estuaries:scales of distribution and relationships with environmental variables[J]. Journal of Experimental Marine Biology and Ecology, 270:1-24.
Feder HM, Jewett SC.2007. Blanchard AL.Southeastern Chukchi Sea (Alaska) macrobenthos[J]. Ploar Biology,30:261-275.
Fujii T.2007. Spatial patterns of benthic macrofauna in relation to environmental variables in an intertidal habitat in the Humber estuary, UK:Developing a tool for estuarine shoreline managemen[J]. Estuarine, Coastal and Shelf Science,75:101-119.
Honkoop PJC, Pearson GB, Lavaleye MSS, et al.2006. Spatial variation of the intertidal sediments and macrozoo-benthic assemblages along Eighty-mile Beach, Northwestern Australia[J]. Journal of Sea Research,55:278-291.
Peter RT, Wookdridge TH.2003. What limits the distribution of subtidal macrobenthos in permanently open and temporarily open/closed South African estuaries? Salinity vs. sediment paricle size[J]. Estuarine,Coastal and Shelf Science,57:225-238.
Spruzen FL, Richardson AMM, Woehler EJ.2008. Spatial variation of intertidal macroinvertebrates and environmental variables in Robbins Passage wetland, NW Tasmania[J]. Hydrobiologia,598:325-342.
Taft OW, Haig SM.2005. The value of agricultural wetlands as invertebrate resources for wintering shorebirds[J].Agriculture, Ecosystems and Environment,110:249-256.
Thrush SF, Hewitt JE, GibbsM, et al.2006.Functional role of large organisms in intertidal communities: community effects and ecosystem function[J]. Ecosystem,9:1029-1040.
Wieking G, Kroncke I.2005. Is benthic trophic structure affected by food quality? The Dogger Bank example[J]. Marine Biology,146:387-400.
鲍毅新,葛宝明,郑祥,等.2006.温州湾天河滩涂大型底栖动物群落分布与季节变化[J].动物学报,52(1)：45-52.
鲍毅新,胡知渊,李欢欢,等.2008.灵昆东滩围垦区内外大型底栖动物季节变化和功能群的比较[J].动物学报,54(3)：416-427.
葛宝明,鲍毅新,程宏毅,等.2008.灵昆岛东滩潮间带大型底栖动物功能群及营养等级构成[J].生态学报,28(10)：4796-4804.
葛宝明,鲍毅新,郑祥.2005.灵昆岛围垦滩涂潮沟大型底栖动物群落生态学研究[J].生态学报,25(3)：446-452.
葛宝明,郑祥,程宏毅,等.2007.灵昆岛围垦滩涂潮沟大型底栖动物群落和物种生态位分析[J].水生生物学报,31(5)：675-681.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报,28(4)：1498-1507.
金亮,蔡立哲,周细平,等.2007.深圳湾北岸泥滩大型底栖动物次级生产力研究[J].台湾海峡,26(3)：413-421.
李欢欢,鲍毅新,胡知渊,等.2007.杭州湾南岸大桥建设区域潮间带大型底栖动物功能群及营养等级的季节动态[J].动物学报,53(6)：1011-1023.
李新正,于海燕,王永强,等.2001.胶州湾大型底栖动物的物种多样性现状[J].生物多样性,9(1)：80-84.
厉红梅,蔡立哲,林丽珠,等.2001.深圳湾潮间带底栖动物群落结构的等级聚类与非度量多维标度测序[J].厦门大学学报,40(3)：35-740.
刘建军,徐艳华.1997.水磨河底栖动物群落结构的生态位分析[J].水生生物学报,21(2)：101-108.
刘奕琳.2006.盐城海滨湿地生态系统的研究[A].博士学位论文.南京林业大学.
马藏允,刘海,王惠卿,等.1997.底栖动物群落结构变化多元变量统计分析[J].中国环境科学,17(4)：297-300.
马克平,刘玉明.1994.生物群落多样性的测度方法I.a多样性的测定方法(下)[J].生物多样 性,2(4)：231-239.
马克平.1994.生物多样性的测度方法.钱迎倩,马克平主编.生物多样性研究的原理与方法[M].北京：中国科学技术出版社,141-165.
任美锷,许廷官,朱季文,等.1986.江苏省海岸带和海涂资源综合调查报告[M].北京：海洋出版社.
王加连,吕士成.2008.江苏省盐城滩涂野生动物资源调查研究[J].四川动物,27(4)：620-625.
徐治国,何岩,闫百兴,等.2007.三江平原典型沼泽湿地植物种群的生态位[J].应用生态学报,18(4)：783-787.
袁兴中,陆健健.2001.长江口岛屿湿地的底栖动物资源研究[J].自然资源学报,16(1)：35-41.
袁兴中,陆健健,刘红.2002.长江口底栖动物功能群分布格局及其变化[J].生态学报,22(12)：2054-2062.
赵永强,曾江宁,高爱根,等.2009.椒江口潮间带大型底栖动物的生态位[J].应用生态学报,20(5)1176-1183.
郑荣泉,张永普,李灿阳,等.2007.乐清湾滩涂大型底栖动物群落结构的时空变化[J].动物学报,53(2)：390-398.
周晓,葛振鸣,施文或,等.2006.长江口九段沙湿地大型底栖动物群落结构的季节变化规律[J].应用生态学报,17(11)：2079-2083.
周晓,葛振鸣,施文或,等.2007.长江口新生湿地大型底栖动物群落时空变化格局[J].生态学杂志,26(3)372-377.
朱晓佳,钦佩.2003.外来种互花米草及米草生态工程[J].海洋科学,27(12)：14-19.
Angradi TR, Hagan SM, Able KW.2001. Vegetation type and the intertidal macro-invertebrate fauna of a brackish marsh:Phragmites vs. Spartina[J]. Wetland,21(9):75-92.
Cordell JR, Simenstad CA, Feist B, et al.1998.Ecological effects of S partina alterniflora invasion of the Littoral flat community in Willapa Bay,Washington[A].Abstracts from the Eighth International Zebra Mussel and Other Nuisance Species Conference, Sacramento California.
Curtis CD, Donald RS.1996. Status, prediction and prevention of introduced cordgrass Spartina spp. Invasion in pacific estuaries, USA[J]. Biological Conservation,78:51-58.
Hedge P, Kriwoken LK.2000. Evidence for effects of Spartina anglica invasion on macrobenthic fauna in Little Swanport estuary, Tasmania[J]. Austral Ecology,25:150-159.
Lana P, Guiss C.1991. Influence of Spartina alterniflora on structure and temporal variability of macrobenthic associations intidal flat of Paranagua Bay,Brazil[J]. Marine Ecolog Progress Series,9(73):231-234.
Luiting VT, Cordell JR, Olson AM,et al.1997.Doex exotic Spartina alterniflora change benthic invertebrate assemblages. In:Pattern K (ed.), Proceedings of the Second International Spartina Conference[A]. Washington State University,Olympia,48-50.
Neira C, Grosholz ED, Levin LA, et al.2006. Mechanisms generating modification of benthos following tidal flat invasion by a Spartina hybrid[J]. Ecological applications,16(17):1391-1404.
Posey MH, Alphin TD, Meyer DL, et al.2003.Benthic communities of common reed Phragmites australis and marsh cordgrass S partina alterniflora marshes in Chesapeake Bay[J]. Marine Ecology-Progress Series, 261:51-61.
Posey MH, Alphin TD, Meyer DL, et al.2003. Benthic communities of common reed Phragmites australis and marsh cordgrass Spartina alterniflora marshes in Chesapeake Bay[J]. Marine Ecology- Progress Series, 261:51-61.
Talley TS, Levin LA.2001. Modification of sediment s and macrofauna by an invasive marsh plant[J]. Biological Invasions,3:51-68.
Zhang RS, Shen YM, Lu LY, et al.2004. Formation of Spartina alterniflora salt marshes on the coast of Jiangsu Province[J].China.Ecol,23(2):95-105.
Zhao YJ, Zeng JN, Chen QZ, et al.2009. Community pattern of macrozoobenthos in distinct habitats with different densities of Spartina alterniflora Loisel[J]. Journal of Natural Resources,24(4):631-638.
Zhou HX, Liu JE, Qin P.2009. Impacts of an alien species(Spartina alterniflora) on the macrobenthos community of Jiangsu coastal inter-tidal ecosystem[J]. Ecological Engineering,35:521-528.
陈忠义,付萃长,王海毅,等.2005.互花米草入侵东滩盐沼对大型底栖无脊椎动物群落的影响[J].湿地科学,3(1)：1-7.
葛宝明,鲍毅新,郑祥.2005.灵昆岛围垦滩涂潮沟大型底栖动物群落生态学研究[J].生态学报,25(3)：446-452.
洪荣标,吕小梅,陈岚,等.2005.九龙江口红树林湿地与米草湿地的底栖生物[J].台湾海峡,24(2)：189-194.
胡知渊,李欢欢,鲍毅新,等.2008.灵昆岛围垦区内外滩涂大型底栖动物生物多样性[J].生态学报,28(4)：1498-1507.
田家怡,申保忠,李建庆,等.2009.黄河三角洲外来入侵物种米草对滩涂底栖动物的影响[J].海洋环境科学,28(6)：687-690.
谢文静,高抒.2009.江苏中部海岸互花米草盐沼底栖动物的组成特征[J].南京大学学报(自然科学),45(4)：553-560.
谢志发,何文珊,刘文亮,等.2008.不同发育时间的互花米草盐沼对大型底栖动物群落的影响[J].生态学杂志,27(1)：63-67.
谢志发,章飞军,刘文亮,等.2007.长江口互花米草生长区大型底栖动物的群落特征(英文)[J].动物学研究.28(2)：167-171.
任美锷,许廷官,朱季文,等.1986.江苏省海岸带和海涂资源综合调查报告[M].北京：海洋出版社.
徐炳声.1999.上海植物志(下卷)[M]上海：上海科学技术文献出版社.
徐晓军,王华,由文辉,等.2006.崇明东滩互花米草群落中底栖动物群落动态的初步研究[J].海洋湖沼通报,2：89-95.
杨泽华,童春富,陆健健.2007.盐沼植物对大型底栖动物群落的影响[J].生态学报,27(11)：4387-4393.
虞蔚岩,李朝晖,华春,等.2009a.江苏盐城东台互花米草滩涂底栖无脊椎动物的多样性分析[J].海洋湖沼通报.1(6)：123-128.
虞蔚岩,李朝晖,黄成,等.2009b.互花米草入侵对盐城东台盐滩多毛纲动物多样性的影响及管理对策[J].上海海洋大学学报,18(1)：47-5.
赵永强,曾江宁,陈全震,等.2009.不同互花米草(Spartina alterniflora Loisel)密度生境中大型底栖动物群落格局[J].自然资源学报,24(4)：631-638.
仲崇信.1985.大米草简史及国外研究概况[A].南京大学学报(米草研究的进展-22年来的研究成果论文集),40(2)：1-30.
周晓,王天厚,葛振鸣,等.2006.长江口九段沙湿地不同生境中大型底栖动物群落结构特征分析[J].生物多样性,14(2)：165-171.
周晓,王天厚,葛振鸣,等.2006.长江口九段沙湿地不同生境中大型底栖动物群落结构特征分析[J].生物多样性,14(2)：165-171.
朱晓佳,钦佩.2003.外来种互花米草及米草生态工程[J].海洋科学,27(12)：14-19.
Battley PF, Piersma T.2005. Body composition and flight ranges of Bar-tailed Godwit (Limosa lapponica Baueri) from New Zealand[J]. Auk,122(3):922-937.
Battley PF, Piersma T, Rogers DI, et al.2004.Do body condition and plumage during fuelling predict northward departure dates of Great knots Calidris tenuirostris from northwest Australia? [J] Ibis,146:46-60.
Danufsky T, Colwell MA.2003.Winter shorebird communities and tidal flat characteristics at Humbodt Bay, Califonia[J]. The Condor,105:117-129.
Gan XJ, Cai YT, Choi CY, et al.2009. Potential impacts of invasive Spartina alterniflora on spring bird communities at Chongming Dongtan, a Chinese wetland of international importance [J].Estuarine,Coastal and Shelf Science,83:211-218.
Howes J, Bakewell D.1989.Shorebird Studies Manual. Kuala Lumpur:AWB Publication.143-147.
Ma ZJ, Li B, Zhao B, et al.2004. Are artificial wetlands good alternatives to natural wetlands for waterbirds? —A case study on Chongming Island, China[J]. Biodiversity and Conservation,13:333-350.
Piersma T, DeGoeij P, Tulp I.1993. An evalution of intertidal feeding habitats from a shorebird perspective towards relevant comparisons between temperate and tropical mudflats Netherlands [J] Journal of Sea Research,31:103-123.
Shepherd PCF, Lank DB.2004. Marine and agricultural habitat preferences of dunlin wintering in British Columbia[J]. Journal of Wildlife Management,68(1):61-73.
Weber LM, Haig SM.1997. Shorebird diet and size selection of nereid polychaetes in South Carolina coastal diked wetlands[J]. Journal of Field Omithology.68(3):358-366
Wilson JR, Barter M.1998. Identification of potentially important staging area for 'long jump' migrant waders in the east Asian-Australian flyway during northward migration[J]. Silt,32:16-26.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M].北京：中国林业出版社.
崔志兴,钱国祯,祝龙彪,等.1985.鸻形目鸟类的食性研究[J]动物学研究(增刊).6(4)：43-51.
葛振鸣,王天厚,施文或,等.2006a.长江口杭州湾鸻形目鸟类群落季节变化和生境选择[J].生态学报,26(1)：40-47.
葛振鸣,王天厚,周晓等.2006b.上海崇明东滩堤内次生人工湿地鸟类冬春季生境选择的因子分析[J].动物学研究,27(2)：144-150.
葛振鸣,周晓,施文或,等.2007.九段沙湿地鸻形目鸟类迁徙季节环境容纳量生态学报[J].27(1)：90-96.
国家环保总局.2003.优先保护17个生物多样性关键地区[J].生态经济,2：78.
金杰锋,刘伯锋,余希,等.2008.福建省兴化湾滨海养殖塘冬季水鸟的栖息地利用[J].动物学杂志,43(6)：17-24.
敬凯.2005.上海崇明东滩鸻鹬类中途停歇生态学研究.复旦大学(博士论文).
李鑫,吴建平,朱成伟,等.2010.卧龙湖自然保护区水鸟迁徙研究[J].野生动物,31(3)：135-138.
刘伯锋.2003.福建沿海湿地鸻鹬类资源调查[J].动物学杂志,38(6)：72-75.
马敬能,菲利普斯,何芬奇.2000.中国鸟类野外手册[J].湖南教育出版社.
唐承佳,陆健健.2002.围垦堤内迁徙鸻鹬群落的生态学特性[J].动物学杂志,37(2)：27-33.
王会,杜进进.1993.射阳盐场湿地禽类资源考察初报[J].动物学杂志,28(4)：21-24.
张国钢,梁伟,刘冬平,等.2005.海南岛越冬水鸟资源状况调查[J].动物学杂志,40(2)：80-85.
朱晶,敬凯,干晓静,等.2007.迁徙停歇期鸻鹬类在崇明东滩潮间带的食物分布[J].生态学报.27(6)：2149-2159.
Barbosa A, Moreno E.1999.Evolution of foraging strategies in shorebirds:an ecomophorlogical approach[J]. Auk,116:712-725.
Boettcher R, Haig SM, Bridges WC.1995. Habitat-related factors affecting the distribution of nonbreeding Ameriean Avocets in Coastal South-Carolina[J]. Condor,97:68-81.
Bryant, DM.1979. Effects of prey density and site character on estuary usage by overwintering waders (Charadrii) [J]. Estuarine and Coastal Marine Science,9:369-384.
Butler RW, Davidson NC, Morrison RIG.2001.Global-scale shorebird distribution in relation to productivity of near-shore ocean waters[J]. Waterbirds,24:224-232.
Davis CA, Smith LM.2001. Foraging strategies and niche dynamics of coexisting shorebirds at stopover sites in the southern Great Plains[J] Auk,118:484-495.
Davis CA,Smith LM.2001.Foraging strategies and niche dynamics of coexisting shorebirds at stopover sites in the southern Great Plains[J].Auk,118:484-495.
Goss-Custard JD.1977. The ecology of the Wash. Ⅲ. Density-related behaviour and the possible effects of a loss of feeding grounds on wading birds (Charadrii) [J]. Journal of Applied Ecology,14:721-739.
Jing K, Ma ZJ, Li B,et al.2007.Foraging strategies involved in habitat use of shorebirds at the intertidal area of Chongming Dongtan,China[J].Ecologiccal Research,22:559-570.
Kalejta B, Hockey PAR.1994. Distribution of Shorebirds at the Berg River estuary, South Africa, in relation to foraging mode, food supply and environmental features[J]. Ibis,136:233-239.
Moreira F.1999. On the use by birds of intertidal areas of the Tagus estuary:implications for management[J]. Aquatic Ecology 33:301-309.
Morrison RIG.1984 Migration system of some new world shorebirds.In:Olla BL(ed)Shorebirds:migration and foraging behavior[M].Plenum Press:New York,125-202.
Nuka T, Norman CP, Kuwabara K, et al.2005.Feeding behavior and effect of prey availability on Sanderling Calidris alba distribution on Kujukuri Beach[J].Omithol. Sci,4:139-146.
Placyk JSJ, Harrington BA.2004 Prey abundance and habitat use by migratory shorebirds at coastal stopover sites in Connecticut[J]. Journal of Field Ornithology,75:223-231.
Ribeiro PD, Iribarne OO, Navarro D, et al.2004. Environmental heterogeneity, spatial segregation of prey, and the utilisation of southwest Atlantic mudflats by migratory shorebirds[J]. Ibis 146:672-683.
Ribeiro PD, Iribarne 00, Navarro D, et al.2004. Environmental heterogeneity, spatial segregation of prey, and the utilisation of southwest Atlantic mudflats by migratory shorebirds [J]. Ibis,146:672-683.
Rogers D I.2002. What determines shorebirds feeding distribution in Roebuck Bay?[C].M Pepping et al.(eds).Intertidal sediments and benthic animals of Roebuck Bay Western Australia. Texel:Nertherland Institute for Sea Research(NIOZ),145-174.
Sanchez MI, Green AJ, Castellanos EM.2006. Spatial and temporal fluctuations in presence and use of chironomid prey by shorebirds in the Odiel saltpans,south-west Spain[J]. Hydrobiologia,567:329-340.
Schoener TW.1965. The evolution of bill size differences among sympatric congeneric species of birds[J].Evolution,19:189-213.
Spruzen FL, Richardson AMM, Woehler EJ.2008. Influence of environmental and prey variables on low tide shorebird habitat use within the Robbins Passage wetlands, Northwest Tasmania[J]. Estuarine, Coastal and Shelf Science,78:122-134.
Zharikov Y, Skilleter GA.2002. Sex-specific intertidal habitat use in subtropically wintering Bar-tailed Godwits[J]. Canadian Journal of Zoology. (Revue Canadienne de Zoologie),80:1918-1929.
Zou FS, Zhang HH, Dahmer T, et al.2008.The effects of benthos and wetland area on shorebird abundance and species richness in coastal mangrove wetlands of Leizhou Peninsula, China [J]. Forest Ecology and Management,255:3813-3818.
陈克林主编.2006.黄渤海湿地与迁徙水鸟进行研究[M].北京：中国林业出版社.
黑龙江省野生动物研究所.1992.黑龙江省鸟类志[M].北京：中国林业出版社.136-165.
孙儒泳.2001.动物生态学原理[M].北京：北京师范大学出版社.
唐仕华,虞快.1996.崇明东滩鸻形目鸟类群落及其食性研究[J].华东师范大学学报(自然科学版)动物学专辑,79-83.
袁兴中,陆健健.2001.长江口潮沟大型底栖动物群落的初步研究[J].动物学研究,22：211-215.
朱书玉,吕卷章,赵长征,等.2000.黄河三角洲国家级自然保护区鸻形目鸟类食性及觅食地的研究[J].山东林业科技,130：10-13.
© 2004-2018 中国地质图书馆版权所有 京ICP备05064691号 京公网安备11010802017129号 地址：北京市海淀区学院路29号 邮编：100083 电话：办公室：(+86 10)66554848；文献借阅、咨询服务、科技查新：66554700 |