经营管理对森林土壤有机碳库影响的研究进展
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摘要
森林土壤有机碳库在全球碳循环及减缓气候变暖中发挥着重要作用。施肥、火烧、采伐、林下植被管理、覆盖等营林措施改变了森林生态系统的生产力,显著影响了森林土壤的碳输入和碳输出。综述了主要营林措施对森林土壤有机碳库的影响及其机制,并探讨今后的研究重点。总体而言,施用有机肥、有机无机肥配施及生物质炭添加均可提高土壤有机碳含量。氮肥对土壤有机碳含量的影响存在增加、降低和无影响3种结果;火烧对土壤有机碳的影响取决于火烧后恢复时间、火烧温度、火烧强度、土层深度等因素;皆伐通过改变土壤温度、含水量、有机碳来源等因素,导致森林土壤有机碳含量下降;而间伐对土壤有机碳的影响则与间伐强度有关;去除林下植被及凋落物加快了土壤有机碳的分解,但林下植被的替代与添加则会提高土壤有机碳含量;覆盖提高了土壤有机碳含量,但导致有机碳稳定性下降。随着研究方法和观测手段的不断发展,今后应深入研究营林措施对土壤碳形态、结构和转化过程的影响;同时,更多关注人为管理和气候变化对森林土壤碳库产生的叠加效应。
Forest soil as an important carbon sink plays a critical role in the global carbon cycle and mitigation of climate warming. Fertilization, fire, cutting, understory management an dmulching have changed the productivity of forest ecosystem, which significantly affects the carbon input and carbon output of forest soil. This paper analyzed the effects of different kinds of forest management practices on soil organic carbon pool and proposed important directions for future research, in order to improve the carbon sequestration capacity of plantation soils through forest management practices. Organic fertilizer, organic-inorganic fertilizer and biochar tended to improve the soil active carbon content. N fertilizers reduced the soil active organic carbon content in N-rich forests, but there was an increase or no significant difference in soil active organic carbon content in N-poor forest. The influence of fire on soil organic carbon pool was determined by time length after fire, fire temperature, fire intensity and soil depth. Clear-cut changed the soil temperature, water content and organic matter sources, and thus reduced the organic carbon storage of forest soil. The effect of thinning on soil organic carbon content was related to the harvesting intensity. Understory weeding enhanced soil temperature which could accelerate the decomposition of soil organic carbon, whereas the replacement and addition of understory vegetation would do the other way around. Mulching of organicresidues in forests helped to increase the soil organic carbon content but decreased its stability. With the development of research theory and technology, the impact of forest management practiceson soil carbon form and structure onsoil carbon form and structure will become the main direction for future research.
Forest soil as an important carbon sink plays a critical role in the global carbon cycle and mitigation of climate warming. Fertilization, fire, cutting, understory management an dmulching have changed the productivity of forest ecosystem, which significantly affects the carbon input and carbon output of forest soil. This paper analyzed the effects of different kinds of forest management practices on soil organic carbon pool and proposed important directions for future research, in order to improve the carbon sequestration capacity of plantation soils through forest management practices. Organic fertilizer, organic-inorganic fertilizer and biochar tended to improve the soil active carbon content. N fertilizers reduced the soil active organic carbon content in N-rich forests, but there was an increase or no significant difference in soil active organic carbon content in N-poor forest. The influence of fire on soil organic carbon pool was determined by time length after fire, fire temperature, fire intensity and soil depth. Clear-cut changed the soil temperature, water content and organic matter sources, and thus reduced the organic carbon storage of forest soil. The effect of thinning on soil organic carbon content was related to the harvesting intensity. Understory weeding enhanced soil temperature which could accelerate the decomposition of soil organic carbon, whereas the replacement and addition of understory vegetation would do the other way around. Mulching of organicresidues in forests helped to increase the soil organic carbon content but decreased its stability. With the development of research theory and technology, the impact of forest management practiceson soil carbon form and structure onsoil carbon form and structure will become the main direction for future research.
引文
[1] LIU Xiang, YANG Tao, WANG Quan, et al. Dynamics of soil carbon and nitrogen stocks after afforestation in arid and semi-arid regions:a meta-analysis[J]. Sci Total Environ, 2018, 618:1658-1664.
[2] KEENAN R J, REAMS G A, ACHARD F, et al. Dynamics of global forest area:results from the FAO global forest resources assessment[J]. For Ecol Manage, 2015, 352(7):9-20.
[3] LAL R. Soil carbon sequestration to mitigate climate change[J]. Geoderma, 2004, 123(1/2):1-22.
[4] PAN Yude, BIRDESY R A, FANG Jinyun, et al. A large and persistent carbon sink in the world’s forests[J]. Science, 2011, 333(6045):988-992.
[5] LAL R. Forest soils and carbon sequestration[J]. For Ecol Manage, 2005, 220(1/3):242-258.
[6] NOORMETS A, EPRON D, DOMEC J C, et al. Effects of forest management on productivity and carbon sequestration:a review and hypothesis[J]. For Ecol Manage, 2015, 355:124-140.
[7]张蛟蛟,李永夫,姜培坤,等.施肥对板栗林土壤活性碳库和温室气体排放的影响[J].植物营养与肥料学报,2013, 19(3):745-752.ZHANG Jiaojiao, LI Yongfu, JIANG Peikun, et al. Effect of fertilization on labile carbon pools and emissions of greenhouse gas in soils of Chinese chestnut stands[J]. Plant Nutri Fert Sci, 2013, 19(3):745-752.
[8]陈雪双,刘娟,姜培坤,等.施肥对山核桃林地土壤N2O排放的影响[J].植物营养与肥料学报, 2014, 20(5):1262-1270.CHEN Xueshuang, LIU Juan, JIANG Peikun, et al. Effects of fertilization on soil N2O flux in Chinese Carya cathayensis stands[J]. Plant Nutr Fert Sci, 2014, 20(5):1262-1270.
[9]姜培坤,徐秋芳.施肥对雷竹林土壤活性有机碳的影响[J].应用生态学报, 2005, 16(2):253-256.JIANG Peikun, XU Qiufang. Effects of fertilization on soil active organic carbon under Phyllostachys praecox stand[J]. Chin J Appl Ecol, 2005, 16(2):253-256.
[10]涂利华,胡庭兴,张健,等.模拟氮沉降对华西雨屏区苦竹林土壤有机碳和养分的影响[J].植物生态学报,2011, 35(2):125-136.TU Lihua, HU Tingxing, ZHANG Jian, et al. Response of soil organic carbon and nutrients to simulated nitrogen deposition in Pleioblastus amarus plantation, rainy area of west China[J]. Chin J Plant Ecol, 2011, 35(2):125-136.
[11]汪金松,赵秀海,张春雨,等.模拟氮沉降对油松林土壤有机碳和全氮的影响[J].北京林业大学学报, 2016, 38(10):88-94.WANG Jinsong, ZHAO Xiuhai, ZHANG Chunyu, et al. Effects of simulated nitrogen deposition on soil organic carbon and total nitrogen content in plantation and natural forests of Pinus tabuliformis[J]. J Beijing For Univ, 2016, 38(10):88-94.
[12] ALLISON S D, CZIMCZIK C I, TRESEDER K K. Microbial activity and soil respiration under nitrogen addition in Alaskan boreal forest[J]. Global Change Biol, 2008, 14(5):1156-1168.
[13] MAGILL A H, ABER J D, CURRIE W S, et al. Ecosystem response to 15 years of chronic nitrogen additions at the harvard forest LTER, Massachusetts, USA[J]. For Ecol Manage, 2004, 196(1):7-28.
[14]郭亮,王庆贵,邢亚娟.森林生态系统土壤碳库对氮沉降的响应研究进展[J].中国农学通报, 2016, 32(32):81-87.GUO Liang, WANG Qinggui, XING Yajuan. Response of forest ecosystem soil carbon pool to nitrogen deposition:a review[J]. Chin Agric Sci Bull, 2016, 32(32):81-87.
[15] NOYCE G L, BASILIKO N, FULTHORPE R, et al. Soil microbial responses over 2 years following biochar addition to a north temperate forest[J]. Biol Fert Soil, 2015, 51(6):649-659.
[16] SUN Liying, LI Lu, CHEN Zhaozhi, et al. Combined effects of nitrogen deposition and biochar application on emissions of N2O, CO2and NH3from agricultural and forest soils[J]. Soil Sci Plant Nutr, 2014, 60(2):254-265.
[17] LIANG Biqing, LEHMANN J, SOHI S P, et al. Black carbon affects the cycling of non-black carbon in soil[J]. Organic Geochem, 2010, 41(2):206-213.
[18]尹艳,刘岩,尹云锋,等.生物质炭添加对杉木人工林土壤原有有机碳矿化的影响[J].应用生态学报, 2018, 29(5):1389-1396.YIN Yan, LIU Yan, YIN Yunfeng, et al. Effects of biochar addition on the mineralization of native soil organic carbon in Cunninghamia lanceolata plantation[J]. Chin J Appl Ecol, 2018, 29(5):1389-1396.
[19] LIU Shuwei, ZHANG Yaojun, ZONG Yajie, et al. Response of soil carbon dioxide fluxes, soil organic carbon and microbial biomass carbon to biochar amendment:a meta-analysis[J]. Global Change Biol, 2016, 8(2):392-406.
[20] PIASZCZYK W, B覵O'NSKA E, LASOTA J. Study on the effect of organic fertilizers on soil organic matter and enzyme activities of soil in forest nursery[J]. Soil Sci Ann, 2017, 68(3):125-131.
[21]姜培坤,徐秋芳,邬奇峰,等.施肥对板栗林土壤养分和生物学性质的影响[J].浙江农林大学学报, 2007, 24(4):445-449.JIANG Peikun, XU Qiufang, WU Qifeng, et al. Effect of fertization on soil properties under Castanea mollissima plantation[J]. J Zhejiang A&F Univ, 2007, 24(4):445-449.
[22]王伟.施肥对梁山慈竹林土壤有机碳和酶活性的影响[D].雅安:四川农业大学, 2012.WANG Wei. The Impact of Fertilization on Organic Carbon and Enzyme Activities of the D. farinosus Chia et H.L.Fung Soil[D]. Ya’an:Sichuan Agricultural University, 2012.
[23]向元彬,周世兴,肖永翔,等.模拟氮沉降和降雨量改变对华西雨屏区常绿阔叶林土壤有机碳的影响[J].生态学报, 2017, 37(14):4686-4695.XIANG Yuanbin, ZHOU Shixing, XIAO Yongxiang, et al. Effects of simulated nitrogen deposition and precipitation changes on soil organic carbon in an evergreen broad-leaved forest that is part of the rainy area of western China[J].Acta Ecol Sin, 2017, 37(14):4686-4695.
[24]郑威.施氮对樟树人工林土壤呼吸的影响[D].长沙:中南林业科技大学, 2013.ZHENG Wei. Effect of Nitrogen Fertilization of Soil Respiration in Cinnamomum camphora Plantation[D]. Changsha:Central South University of Forestry and Technology, 2013.
[25]张明江,龙健,李娟,等.生物质炭对茂兰喀斯特森林土壤呼吸与有机碳的影响[J].土壤, 2018, 50(2):333-340.ZHANG Mingjiang, LONG Jian, LI Juan, et al. Effects of biochars on soil respiration and SOC content of origin forest soil in Maolan Karst Area of Guizhou, China[J]. Soils, 2018, 50(2):333-340.
[26]张岩,张令珍,徐美丽,等.太岳山油松林土壤活性碳和微生物特性随外源有机物的变化规律[J].福建农林大学学报, 2017, 46(3):284-292.ZHANG Yan, ZHANG Lingzhen, XU Meili, et al. Variations in labile soil carbon and microbial activity by organic matter input in Pinus forest in Taiyue mountain[J]. J Fujian A&F Univ, 2017, 46(3):284-292.
[27] FLORENT M, FIELD C B. Fire history and the global carbon budget:a 1°×1°fire history reconstruction for the 20th century[J]. Global Change Biol, 2010, 11(3):398-420.
[28] CHEN H Y H, SHRESTHA B M. Stand age, fire and clear cutting affect soil organic carbon and aggregation of mineral soils in boreal forests[J]. Soil Biol Biochem, 2012, 50(5):149-157.
[29] HATTEN J A, ZABOWSKI D, OGDEN A, et al. Soil organic matter in a ponderosa pine forest with varying seasons and intervals of prescribed burn[J]. For Ecol Manage, 2008, 255(7):2555-2565.
[30]薛立,陈红跃,杨振意,等.火灾对马尾松林地土壤特性的影响[J].生态学报, 2011, 31(22):6824-6831.XUE Li, CHEN Hongyue, YANG Zhenyi, et al. The effect of fire on soil properties in a Pinus massoniana stand[J].Acta Ecol Sin, 2011, 31(22):6824-6831.
[31] WANG Qingkui, ZHONG Micai, WANG Silong. A meta-analysis on the response of microbial biomass, dissolved organic matter, respiration, and N mineralization in mineral soil to fire in forest ecosystems[J]. For Ecol Manage,2012, 271:91-97.
[32]孙龙,赵俊,胡海清.中度火干扰对白桦落叶松混交林土壤理化性质的影响[J].林业科学, 2011, 47(2):103-110.SUN Long, ZHAO Jun, HU Haiqing. Effect of moderate fire disturbance on soil physical and chemical properties of Betula platyphylla-Larix gmelinii mixed forest[J]. Sci Silv Sin, 2011, 47(2):103-110.
[33] JOHNSON D W, CURTIS P S. Effects of forest management on soil C and N storage:meta analysis[J]. For Ecol Manage, 2001, 140(2/3):227-238.
[34] K魻STER E, K魻STER K, BERNINGER F, et al. Carbon dioxide, methane and nitrous oxide fluxes from podzols of a fire chronosequence in the boreal forests in V覿rri觟, Finnish Lapland[J]. Geoderma, 2015, 5:181-187.
[35]郭剑芬,杨玉盛,陈光水,等.火烧对森林土壤有机碳的影响研究进展[J].生态学报, 2015, 35(9):2800-2809.GUO Jianfen, YANG Yusheng, CHEN Guangshui, et al. A review of effects of fire on soil organic carbon in forests[J]. Acta Ecol Sin, 2015, 35(9):2800-2809.
[36]许鹏波,屈明,薛立.火对森林土壤的影响[J].生态学杂志, 2013, 32(6):1596-1606.XU Pengbo, QU Ming, XUE Li. Effects of forest fire on forest soils[J]. Chin J Ecol, 2013, 32(6):1596-1606.
[37] ALMENDROS G, MART魱N F, GONZ譧LEZVILA F J. Effects of fire on humic and lipid fractions in a Dystric Xerochrept in Spain[J]. Geoderma, 1988, 42(2):115-127.
[38] GONZ譧LEZ-P魪REZ J A, GONZ譧LEZVILA F J, ALMENDROS G. The effect of fire on soil organic matter:a review[J]. Environ Int, 2004, 30(6):855-870.
[39] ALMENDROS G, GONZ譧LEZVILA F J, MARTIN F. Fire-induced transformation of soil organic matter from an oak forest:an experimental approach to the effects of fire on humic substances[J]. Soil Sci, 1990, 149(3):158-168.
[40] ECKMEIER E, GERLACH R, SKJEMSTAD J O, et al. Minor changes in soil organic carbon and charcoal concentrations detected in a temperate deciduous forest a year after an experimental slash-and-burn[J]. Biogeosciences, 2007,4(2):377-383.
[41]吴君君,杨智杰,刘小飞,等.火烧和保留采伐剩余物对土壤有机碳矿化的影响[J].土壤学报, 2015, 52(1):203-211.WU Junjun, YANG Zhijie LIU Xiaofei, et al. Effect of prescribed burning and reservation of logging residues on soil organic carbon mineralization[J]. Acta Pedol Sin, 2015, 52(1):203-211.
[42] TINOCO P, ALMENDROS G, SANZ J, et al. Molecular descriptors of the effect of fire on soils under pine forest in two continental Mediterranean soils[J]. Organic Geochem, 2006, 37(12):1995-2018.
[43] CERTINI G, NOCENTINI C, KNICKER H, et al. Wildfire effects on soil organic matter quantity and quality in two fire-prone Mediterranean pine forests[J]. Geoderma, 2011, 167/168(8):148-155.
[44]杨新芳,鲍雪莲,胡国庆,等.大兴安岭不同火烧年限森林凋落物和土壤C, N, P化学计量特征[J].应用生态学报, 2016, 27(5):1359-1367.YANG Xinfang, BAO Xuelian, HU Guoqing, et al. C∶N∶P stoichiometry characteristics of litter and soil of forests in Great Xing’an Mountains with different fire years[J]. Chin J Appl Ecol, 2016, 27(5):1359-1367.
[45] PAULLIMOGES E, BLACK T A, CHRISTEN A, et al. Effect of clearcut harvesting on the carbon balance of a Douglas-fir forest[J]. Agric For Meteorol, 2015, 203:30-42.
[46] JURGENSEN M E, TARPEY R, PICKENS J, et al. Long term effect of silvicultural thinnings on soil carbon and nitrogen pools[J]. Soil Sci Soc Am J, 2012, 76(4):1418-1423.
[47]闫美芳,张新时,江源,等.主要管理措施对人工林土壤碳的影响[J].生态学杂志, 2010, 29(11):2265-2271.YAN Meifang, ZHANG Xinshi, JIANG Yuan, et al. Effects of management practices on forest plantation soil carbon:a review[J]. Chin J Ecol, 2010, 29(11):2265-2271.
[48] YASHIRO Y, WAN R K, OKUDA T, et al. The effects of logging on soil greenhouse gas(CO2, CH4, N2O)flux in a tropical rain forest, Peninsular malaysia[J]. Agric For Meteorol, 2008, 48(5):799-806.
[49] ZHOU Decheng, ZHAO Shuqing, LIU Shuguang, et al. A meta-analysis on the impacts of partial cutting on forest structure and carbon storage[J]. Biogeosciences, 2013, 10(1):787-813.
[50] YANG A R, SON Y W, NOH N J, et al. Effect of thinning on carbon storage in soil, forest floor and coarse woody debris of Pinus densiflora stands with different stand ages in Gangwon-do, central Korea[J]. For Sci Technol, 2011,7(1):30-37.
[51] SON Y W, YANG A R, CHUNG H, et al. Effect of thinning on soil and forest carbon storage of Quercus stands in Gangwon Province, central Korea[J]. Jpn For Soc Congr, 2011, 122:749-749.
[52]翟凯燕,马婷瑶,金雪梅,等.间伐对马尾松人工林土壤活性有机碳的影响[J].生态学杂志, 2017, 36(3):609-615.ZHAI Kaiyan, MA Tingyao, JIN Xuemei, et al. Effects of thinning intensity on soil active organic carbon in Pinus massoniana plantation[J]. Chin J Ecol, 2017, 36(3):609-615.
[53]郝凯婕.抚育强度对云杉林土壤有机碳含量的影响[J].山西林业科技, 2014, 43(3):16-18.HAO Kaijie. Impact of thinning scheme on soil organic carbon in spruce forest[J]. Shanxi For Sci Technol, 2014, 43(3):16-18.
[54]雷蕾,肖文发.采伐对森林土壤碳库影响的不确定性[J].林业科学研究, 2015, 28(6):892-899.LEI Lei, XIAO Wenfa. Uncertainty effect of forest harvest on soil carbon pool:a review[J]. For Res, 2015, 28(6):892-899.
[55] HU Zhenhong, HE Zongming, HUANG Zhiqun, et al. Effects of harvest residue management on soil carbon and nitrogen processes in a Chinese fir plantation[J]. For Ecol Manage, 2014, 326:163-170.
[56] SMOLANDER A, KITUNEN V, KUKKOLA M, et al. Response of soil organic layer characteristics to logging residues in three scots pine thinning stands[J]. Soil Biol Biochem, 2013, 66:51-59.
[57]方晰,田大伦,项文化.不同经营方式对杉木林采伐迹地土壤C储量的影响[J].中南林业科技大学学报, 2004,24(1):1-5.FANG Xi, TIAN Dalun, XIANG Wenhua. Effects of different management patterns on soil carbon storage of the deforested lands in Chinese fir plantation[J]. J Central South Univ For Technol, 2004, 24(1):1-5.
[58] NAVE L E, VANCE E D, SWANSTON C W, et al. Harvest impacts on soil carbon storage in temperate forests[J].For Ecol Manage, 2010, 259(5):857-866.
[59]郑永,刘小飞,郑蔚,等.应用高频观测探讨不同森林经营方式下矿质土壤呼吸的昼夜动态特征[J].生态学报,2017, 37(1):93-101.ZHENG Yong, LIU Xiaofei, ZHENG Wei, et al. High-frequency analysis of the diel patterns of mineral soil respiration under different forest managemen[J]. Acta Ecol Sin, 2017, 37(1):93-101.
[60] VANGUELOVA E I, PITMAN R, BENHAM S, et al. Impact of tree stump harvesting on soil carbon and nutrients and second rotation tree growth in mid-Wales, UK[J]. Open J For, 2017, 7(1):58-77.
[61]王文杰,张文天,安静,等.落叶松人工林、皆伐迹地及农田土壤碳及肥力的差异[J].林业科学, 2013, 49(9):79-88.WANG Wenjie, ZHANG Wentian, AN Jing, et al. Variation of soil carbons and fertilities in larch plantation land,clear-cut site and farmland in northeast China[J]. Sci Silv Sin, 2013, 49(9):79-88.
[62]丁波,丁贵杰,李先周,等.短期间伐对杉木人工林生态系碳储量的影响[J].中南林业科技大学学报, 2016, 36(8):66-71.DING Bo, DING Guijie, LI Xianzhou, et al. Effects of short term thinning on the carbon storage in Cunninghamia lanceolata plantation ecosystem[J]. J Central South Univer For Technol, 2016, 36(8):66-71.
[63] SETTINERI G, MALLAMACI C, MITROVI'C M, et al. Effects of different thinning intensities on soil carbon storage in Pinus laricio forest of Apennine South Italy[J]. Eur J For Res, 2018, 137(2):131-141.
[64]凌宁.间伐强度对杉木人工林土壤呼吸的影响[D].南京:南京林业大学, 2012.LING Ning. The Impace of Different Thinning Tensity on Soil Respiration in the Chinese fir Plantation[D]. Nanjing:Nanjing Forestry University, 2012.
[65] PAN Ping, ZHAO Fang, NING Jinkui, et al. Impact of understory vegetation on soil carbon and nitrogen dynamic in aerially seeded Pinus massoniana plantations[J]. PLoS One, 2018 1 3(1):e0191952. doi:10.1371/journal.pone.0191952.
[66] WU Jianping, LIU Zhanfeng, CHEN Dima, et al. Understory plants can make substantial contributions to soil respiration:evidence from two subtropical plantations[J]. Soil Biol Biochem, 2011, 43(11):2355-2357.
[67] WANG Xiaoling, ZHAO Jie, WU Jianping, et al. Impacts of understory species removal and/or addition on soil respiration in a mixed forest plantation with native species in southern China[J]. For Ecol Manage, 2011, 261(6):1053-1060.
[68] KUME A, SATOMURA T, TSUBOI N. Effects of understory vegetation on the ecophysiological characteristics of an overstory pine, Pinus densiflora[J]. For Ecol Manage, 2003, 176(1):195-203.
[69] LI Yongfu, JIANG Peikun, CHANG S X, et al. Organic mulch and fertilization affect soil carbon pools and forms under intensively managed bamboo(Phyllostachys praecox)forests in southeast China[J]. J Soils Sediment, 2010, 10(4):739-747.
[70] ZHANG Jiaojiao, LI Yongfu, CHANG S X, et al. Understory vegetation management affected greenhouse gas emissions and labile organic carbon pools in an intensively managed Chinese chestnut plantation[J]. Plant Soil, 2014, 376(1/2):363-375.
[71]吴亚丛,李正才,程彩芳,等.林下植被抚育对樟树人工林土壤活性有机碳库的影响[J].应用生态学报, 2013,24(12):3341-3346.WU Yachong, LI Zhengcai, CHENG Caifang, et al. Effects of understory removal on soil labile organic carbon pool in a Cinnamomum camphora plantation[J]. Chin J Appl Ecol, 2013, 24(12):3341-3346.
[72]李海防,张杏锋.剔除灌草和添加翅荚决明对厚荚相思林土壤温室气体排放的影响[J].应用生态学报, 2010,21(3):563-568.LI Haifang, ZHANG Xingfeng. Soil greenhouse gases emission from an Acacia crassicarpa plantation under effects of understory removal and Cassia alata addition[J]. Chin J Appl Ecol, 2010, 21(3):563-568.
[73] WANG Faming, ZOU Bi, LI Haifang. The effect of understory removal on microclimate and soil properties in two subtropical lumber plantations[J]. J For Res, 2014, 19(1):238-243.
[74] ZHANG Jiaojiao, LI Yongfu, CHANG S X, et al. Understory management and fertilization affected soil greenhouse gas emissions and labile organic carbon pools in a Chinese chestnut plantation[J]. For Ecol Manage, 2015, 337:126-134.
[75] CHATTERJEE S, BANDYOPADHYAY K K, PRADHAN S, et al. Effects of irrigation, crop residue mulch and nitrogen management in maize(Zea mays L.)on soil carbon pools in a sandy loam soil of indo-gangetic plain region[J].Catena, 2018, 165:207-216.
[76] HUANG Zhiqun, XU Zhihong, CHEN Chengrong, et al. Changes in soil carbon during the establishment of a hardwood plantation in subtropical Australia[J]. For Ecol Manage, 2008, 254(1):46-55.
[77]商素云,李永夫,姜培坤,等.天然灌木林改造成板栗林对土壤碳库和氮库的影响[J].应用生态学报, 2012, 23(3):659-665.SHANG Suyun, LI Yongfu, JIANG Peikun, et al. Effects of the conversion from native shrub forest to Chinese chestnut plantation on soil carbon and nitrogen pools[J]. Chin J Appl Ecol, 2012, 23(3):659-665.
[78] MATHERS N J, XU Zhihong. Solid-state13C NMR spectroscopy:characterization of soil organic matter under two contrasting residue management regimes in a 2-year-old pine plantation of subtropical Australia[J]. Geoderma,2003, 114(1/2):19-31.
[79] HUANG Zhiqun, XU Zhihong, CHEN Chengrong. Effect of mulching on labile soil organic matter pools, microbial community functional diversity and nitrogen transformations in two hardwood plantations of subtropical Australia[J].Appl Soil Ecol, 2008, 40(2):229-239.
[80]赵睿宇,李正才,王斌,等.毛竹林地表覆盖年限对土壤有机碳的影响[J].植物生态学报, 2017, 41(4):418-429.ZHAO Ruiyu, LI Zhengcai, WANG Bin, et al. Duration of mulching caused variable pools of labile organic carbon in a Phyllostachys edulis plantation[J]. Chin J Plant Ecol, 2017, 41(4):418-429.
[81]王玉娟.油茶林地覆盖和间种对土壤理化性质及幼林生长的影响[D].长沙:中南林业科技大学, 2009.WANG Yujian. The Effect of Covering and Interplanting on the Soil Physical, Chemical Properties and Growth Condition of Camellia oleifera Young Forest[D]. Changsha:Central South University of Forestry and Technology, 2009.
[82] YOUKHANA A, IDOL T. Tree pruning mulch increases soil C and N in a shaded coffee agroecosystem in Hawaii[J]. Soil Biol Biochem, 2009, 41(12):2527-2534.
[83]郭子武,王为宇,杨清平,等.林地覆盖对雷竹林土壤碳氮磷化学计量特征的影响[J].广西植物, 2013, 33(5):627-632.GUO Ziwu, WANG Weiyu, YANG Qingping, et al. Effect of mulching management on stoichiometry of soil C, N and P in Phyllostachys praecox plantations[J]. Guihaia, 2013, 33(5):627-632.
[84]孙萌,刘洋,李寒,等.有机物覆盖对核桃园土壤有机碳库及酶活性的影响[J].植物营养与肥料学报, 2018, 24(1):270-278.SUN Meng, LIU Yang, LI Han, et al. Effects of organic mulching on soil organic carbon pool and soil enzyme activity in walnut orchard[J]. J Plant Nutr Fert, 2018, 24(1):270-278.
[2] KEENAN R J, REAMS G A, ACHARD F, et al. Dynamics of global forest area:results from the FAO global forest resources assessment[J]. For Ecol Manage, 2015, 352(7):9-20.
[3] LAL R. Soil carbon sequestration to mitigate climate change[J]. Geoderma, 2004, 123(1/2):1-22.
[4] PAN Yude, BIRDESY R A, FANG Jinyun, et al. A large and persistent carbon sink in the world’s forests[J]. Science, 2011, 333(6045):988-992.
[5] LAL R. Forest soils and carbon sequestration[J]. For Ecol Manage, 2005, 220(1/3):242-258.
[6] NOORMETS A, EPRON D, DOMEC J C, et al. Effects of forest management on productivity and carbon sequestration:a review and hypothesis[J]. For Ecol Manage, 2015, 355:124-140.
[7]张蛟蛟,李永夫,姜培坤,等.施肥对板栗林土壤活性碳库和温室气体排放的影响[J].植物营养与肥料学报,2013, 19(3):745-752.ZHANG Jiaojiao, LI Yongfu, JIANG Peikun, et al. Effect of fertilization on labile carbon pools and emissions of greenhouse gas in soils of Chinese chestnut stands[J]. Plant Nutri Fert Sci, 2013, 19(3):745-752.
[8]陈雪双,刘娟,姜培坤,等.施肥对山核桃林地土壤N2O排放的影响[J].植物营养与肥料学报, 2014, 20(5):1262-1270.CHEN Xueshuang, LIU Juan, JIANG Peikun, et al. Effects of fertilization on soil N2O flux in Chinese Carya cathayensis stands[J]. Plant Nutr Fert Sci, 2014, 20(5):1262-1270.
[9]姜培坤,徐秋芳.施肥对雷竹林土壤活性有机碳的影响[J].应用生态学报, 2005, 16(2):253-256.JIANG Peikun, XU Qiufang. Effects of fertilization on soil active organic carbon under Phyllostachys praecox stand[J]. Chin J Appl Ecol, 2005, 16(2):253-256.
[10]涂利华,胡庭兴,张健,等.模拟氮沉降对华西雨屏区苦竹林土壤有机碳和养分的影响[J].植物生态学报,2011, 35(2):125-136.TU Lihua, HU Tingxing, ZHANG Jian, et al. Response of soil organic carbon and nutrients to simulated nitrogen deposition in Pleioblastus amarus plantation, rainy area of west China[J]. Chin J Plant Ecol, 2011, 35(2):125-136.
[11]汪金松,赵秀海,张春雨,等.模拟氮沉降对油松林土壤有机碳和全氮的影响[J].北京林业大学学报, 2016, 38(10):88-94.WANG Jinsong, ZHAO Xiuhai, ZHANG Chunyu, et al. Effects of simulated nitrogen deposition on soil organic carbon and total nitrogen content in plantation and natural forests of Pinus tabuliformis[J]. J Beijing For Univ, 2016, 38(10):88-94.
[12] ALLISON S D, CZIMCZIK C I, TRESEDER K K. Microbial activity and soil respiration under nitrogen addition in Alaskan boreal forest[J]. Global Change Biol, 2008, 14(5):1156-1168.
[13] MAGILL A H, ABER J D, CURRIE W S, et al. Ecosystem response to 15 years of chronic nitrogen additions at the harvard forest LTER, Massachusetts, USA[J]. For Ecol Manage, 2004, 196(1):7-28.
[14]郭亮,王庆贵,邢亚娟.森林生态系统土壤碳库对氮沉降的响应研究进展[J].中国农学通报, 2016, 32(32):81-87.GUO Liang, WANG Qinggui, XING Yajuan. Response of forest ecosystem soil carbon pool to nitrogen deposition:a review[J]. Chin Agric Sci Bull, 2016, 32(32):81-87.
[15] NOYCE G L, BASILIKO N, FULTHORPE R, et al. Soil microbial responses over 2 years following biochar addition to a north temperate forest[J]. Biol Fert Soil, 2015, 51(6):649-659.
[16] SUN Liying, LI Lu, CHEN Zhaozhi, et al. Combined effects of nitrogen deposition and biochar application on emissions of N2O, CO2and NH3from agricultural and forest soils[J]. Soil Sci Plant Nutr, 2014, 60(2):254-265.
[17] LIANG Biqing, LEHMANN J, SOHI S P, et al. Black carbon affects the cycling of non-black carbon in soil[J]. Organic Geochem, 2010, 41(2):206-213.
[18]尹艳,刘岩,尹云锋,等.生物质炭添加对杉木人工林土壤原有有机碳矿化的影响[J].应用生态学报, 2018, 29(5):1389-1396.YIN Yan, LIU Yan, YIN Yunfeng, et al. Effects of biochar addition on the mineralization of native soil organic carbon in Cunninghamia lanceolata plantation[J]. Chin J Appl Ecol, 2018, 29(5):1389-1396.
[19] LIU Shuwei, ZHANG Yaojun, ZONG Yajie, et al. Response of soil carbon dioxide fluxes, soil organic carbon and microbial biomass carbon to biochar amendment:a meta-analysis[J]. Global Change Biol, 2016, 8(2):392-406.
[20] PIASZCZYK W, B覵O'NSKA E, LASOTA J. Study on the effect of organic fertilizers on soil organic matter and enzyme activities of soil in forest nursery[J]. Soil Sci Ann, 2017, 68(3):125-131.
[21]姜培坤,徐秋芳,邬奇峰,等.施肥对板栗林土壤养分和生物学性质的影响[J].浙江农林大学学报, 2007, 24(4):445-449.JIANG Peikun, XU Qiufang, WU Qifeng, et al. Effect of fertization on soil properties under Castanea mollissima plantation[J]. J Zhejiang A&F Univ, 2007, 24(4):445-449.
[22]王伟.施肥对梁山慈竹林土壤有机碳和酶活性的影响[D].雅安:四川农业大学, 2012.WANG Wei. The Impact of Fertilization on Organic Carbon and Enzyme Activities of the D. farinosus Chia et H.L.Fung Soil[D]. Ya’an:Sichuan Agricultural University, 2012.
[23]向元彬,周世兴,肖永翔,等.模拟氮沉降和降雨量改变对华西雨屏区常绿阔叶林土壤有机碳的影响[J].生态学报, 2017, 37(14):4686-4695.XIANG Yuanbin, ZHOU Shixing, XIAO Yongxiang, et al. Effects of simulated nitrogen deposition and precipitation changes on soil organic carbon in an evergreen broad-leaved forest that is part of the rainy area of western China[J].Acta Ecol Sin, 2017, 37(14):4686-4695.
[24]郑威.施氮对樟树人工林土壤呼吸的影响[D].长沙:中南林业科技大学, 2013.ZHENG Wei. Effect of Nitrogen Fertilization of Soil Respiration in Cinnamomum camphora Plantation[D]. Changsha:Central South University of Forestry and Technology, 2013.
[25]张明江,龙健,李娟,等.生物质炭对茂兰喀斯特森林土壤呼吸与有机碳的影响[J].土壤, 2018, 50(2):333-340.ZHANG Mingjiang, LONG Jian, LI Juan, et al. Effects of biochars on soil respiration and SOC content of origin forest soil in Maolan Karst Area of Guizhou, China[J]. Soils, 2018, 50(2):333-340.
[26]张岩,张令珍,徐美丽,等.太岳山油松林土壤活性碳和微生物特性随外源有机物的变化规律[J].福建农林大学学报, 2017, 46(3):284-292.ZHANG Yan, ZHANG Lingzhen, XU Meili, et al. Variations in labile soil carbon and microbial activity by organic matter input in Pinus forest in Taiyue mountain[J]. J Fujian A&F Univ, 2017, 46(3):284-292.
[27] FLORENT M, FIELD C B. Fire history and the global carbon budget:a 1°×1°fire history reconstruction for the 20th century[J]. Global Change Biol, 2010, 11(3):398-420.
[28] CHEN H Y H, SHRESTHA B M. Stand age, fire and clear cutting affect soil organic carbon and aggregation of mineral soils in boreal forests[J]. Soil Biol Biochem, 2012, 50(5):149-157.
[29] HATTEN J A, ZABOWSKI D, OGDEN A, et al. Soil organic matter in a ponderosa pine forest with varying seasons and intervals of prescribed burn[J]. For Ecol Manage, 2008, 255(7):2555-2565.
[30]薛立,陈红跃,杨振意,等.火灾对马尾松林地土壤特性的影响[J].生态学报, 2011, 31(22):6824-6831.XUE Li, CHEN Hongyue, YANG Zhenyi, et al. The effect of fire on soil properties in a Pinus massoniana stand[J].Acta Ecol Sin, 2011, 31(22):6824-6831.
[31] WANG Qingkui, ZHONG Micai, WANG Silong. A meta-analysis on the response of microbial biomass, dissolved organic matter, respiration, and N mineralization in mineral soil to fire in forest ecosystems[J]. For Ecol Manage,2012, 271:91-97.
[32]孙龙,赵俊,胡海清.中度火干扰对白桦落叶松混交林土壤理化性质的影响[J].林业科学, 2011, 47(2):103-110.SUN Long, ZHAO Jun, HU Haiqing. Effect of moderate fire disturbance on soil physical and chemical properties of Betula platyphylla-Larix gmelinii mixed forest[J]. Sci Silv Sin, 2011, 47(2):103-110.
[33] JOHNSON D W, CURTIS P S. Effects of forest management on soil C and N storage:meta analysis[J]. For Ecol Manage, 2001, 140(2/3):227-238.
[34] K魻STER E, K魻STER K, BERNINGER F, et al. Carbon dioxide, methane and nitrous oxide fluxes from podzols of a fire chronosequence in the boreal forests in V覿rri觟, Finnish Lapland[J]. Geoderma, 2015, 5:181-187.
[35]郭剑芬,杨玉盛,陈光水,等.火烧对森林土壤有机碳的影响研究进展[J].生态学报, 2015, 35(9):2800-2809.GUO Jianfen, YANG Yusheng, CHEN Guangshui, et al. A review of effects of fire on soil organic carbon in forests[J]. Acta Ecol Sin, 2015, 35(9):2800-2809.
[36]许鹏波,屈明,薛立.火对森林土壤的影响[J].生态学杂志, 2013, 32(6):1596-1606.XU Pengbo, QU Ming, XUE Li. Effects of forest fire on forest soils[J]. Chin J Ecol, 2013, 32(6):1596-1606.
[37] ALMENDROS G, MART魱N F, GONZ譧LEZVILA F J. Effects of fire on humic and lipid fractions in a Dystric Xerochrept in Spain[J]. Geoderma, 1988, 42(2):115-127.
[38] GONZ譧LEZ-P魪REZ J A, GONZ譧LEZVILA F J, ALMENDROS G. The effect of fire on soil organic matter:a review[J]. Environ Int, 2004, 30(6):855-870.
[39] ALMENDROS G, GONZ譧LEZVILA F J, MARTIN F. Fire-induced transformation of soil organic matter from an oak forest:an experimental approach to the effects of fire on humic substances[J]. Soil Sci, 1990, 149(3):158-168.
[40] ECKMEIER E, GERLACH R, SKJEMSTAD J O, et al. Minor changes in soil organic carbon and charcoal concentrations detected in a temperate deciduous forest a year after an experimental slash-and-burn[J]. Biogeosciences, 2007,4(2):377-383.
[41]吴君君,杨智杰,刘小飞,等.火烧和保留采伐剩余物对土壤有机碳矿化的影响[J].土壤学报, 2015, 52(1):203-211.WU Junjun, YANG Zhijie LIU Xiaofei, et al. Effect of prescribed burning and reservation of logging residues on soil organic carbon mineralization[J]. Acta Pedol Sin, 2015, 52(1):203-211.
[42] TINOCO P, ALMENDROS G, SANZ J, et al. Molecular descriptors of the effect of fire on soils under pine forest in two continental Mediterranean soils[J]. Organic Geochem, 2006, 37(12):1995-2018.
[43] CERTINI G, NOCENTINI C, KNICKER H, et al. Wildfire effects on soil organic matter quantity and quality in two fire-prone Mediterranean pine forests[J]. Geoderma, 2011, 167/168(8):148-155.
[44]杨新芳,鲍雪莲,胡国庆,等.大兴安岭不同火烧年限森林凋落物和土壤C, N, P化学计量特征[J].应用生态学报, 2016, 27(5):1359-1367.YANG Xinfang, BAO Xuelian, HU Guoqing, et al. C∶N∶P stoichiometry characteristics of litter and soil of forests in Great Xing’an Mountains with different fire years[J]. Chin J Appl Ecol, 2016, 27(5):1359-1367.
[45] PAULLIMOGES E, BLACK T A, CHRISTEN A, et al. Effect of clearcut harvesting on the carbon balance of a Douglas-fir forest[J]. Agric For Meteorol, 2015, 203:30-42.
[46] JURGENSEN M E, TARPEY R, PICKENS J, et al. Long term effect of silvicultural thinnings on soil carbon and nitrogen pools[J]. Soil Sci Soc Am J, 2012, 76(4):1418-1423.
[47]闫美芳,张新时,江源,等.主要管理措施对人工林土壤碳的影响[J].生态学杂志, 2010, 29(11):2265-2271.YAN Meifang, ZHANG Xinshi, JIANG Yuan, et al. Effects of management practices on forest plantation soil carbon:a review[J]. Chin J Ecol, 2010, 29(11):2265-2271.
[48] YASHIRO Y, WAN R K, OKUDA T, et al. The effects of logging on soil greenhouse gas(CO2, CH4, N2O)flux in a tropical rain forest, Peninsular malaysia[J]. Agric For Meteorol, 2008, 48(5):799-806.
[49] ZHOU Decheng, ZHAO Shuqing, LIU Shuguang, et al. A meta-analysis on the impacts of partial cutting on forest structure and carbon storage[J]. Biogeosciences, 2013, 10(1):787-813.
[50] YANG A R, SON Y W, NOH N J, et al. Effect of thinning on carbon storage in soil, forest floor and coarse woody debris of Pinus densiflora stands with different stand ages in Gangwon-do, central Korea[J]. For Sci Technol, 2011,7(1):30-37.
[51] SON Y W, YANG A R, CHUNG H, et al. Effect of thinning on soil and forest carbon storage of Quercus stands in Gangwon Province, central Korea[J]. Jpn For Soc Congr, 2011, 122:749-749.
[52]翟凯燕,马婷瑶,金雪梅,等.间伐对马尾松人工林土壤活性有机碳的影响[J].生态学杂志, 2017, 36(3):609-615.ZHAI Kaiyan, MA Tingyao, JIN Xuemei, et al. Effects of thinning intensity on soil active organic carbon in Pinus massoniana plantation[J]. Chin J Ecol, 2017, 36(3):609-615.
[53]郝凯婕.抚育强度对云杉林土壤有机碳含量的影响[J].山西林业科技, 2014, 43(3):16-18.HAO Kaijie. Impact of thinning scheme on soil organic carbon in spruce forest[J]. Shanxi For Sci Technol, 2014, 43(3):16-18.
[54]雷蕾,肖文发.采伐对森林土壤碳库影响的不确定性[J].林业科学研究, 2015, 28(6):892-899.LEI Lei, XIAO Wenfa. Uncertainty effect of forest harvest on soil carbon pool:a review[J]. For Res, 2015, 28(6):892-899.
[55] HU Zhenhong, HE Zongming, HUANG Zhiqun, et al. Effects of harvest residue management on soil carbon and nitrogen processes in a Chinese fir plantation[J]. For Ecol Manage, 2014, 326:163-170.
[56] SMOLANDER A, KITUNEN V, KUKKOLA M, et al. Response of soil organic layer characteristics to logging residues in three scots pine thinning stands[J]. Soil Biol Biochem, 2013, 66:51-59.
[57]方晰,田大伦,项文化.不同经营方式对杉木林采伐迹地土壤C储量的影响[J].中南林业科技大学学报, 2004,24(1):1-5.FANG Xi, TIAN Dalun, XIANG Wenhua. Effects of different management patterns on soil carbon storage of the deforested lands in Chinese fir plantation[J]. J Central South Univ For Technol, 2004, 24(1):1-5.
[58] NAVE L E, VANCE E D, SWANSTON C W, et al. Harvest impacts on soil carbon storage in temperate forests[J].For Ecol Manage, 2010, 259(5):857-866.
[59]郑永,刘小飞,郑蔚,等.应用高频观测探讨不同森林经营方式下矿质土壤呼吸的昼夜动态特征[J].生态学报,2017, 37(1):93-101.ZHENG Yong, LIU Xiaofei, ZHENG Wei, et al. High-frequency analysis of the diel patterns of mineral soil respiration under different forest managemen[J]. Acta Ecol Sin, 2017, 37(1):93-101.
[60] VANGUELOVA E I, PITMAN R, BENHAM S, et al. Impact of tree stump harvesting on soil carbon and nutrients and second rotation tree growth in mid-Wales, UK[J]. Open J For, 2017, 7(1):58-77.
[61]王文杰,张文天,安静,等.落叶松人工林、皆伐迹地及农田土壤碳及肥力的差异[J].林业科学, 2013, 49(9):79-88.WANG Wenjie, ZHANG Wentian, AN Jing, et al. Variation of soil carbons and fertilities in larch plantation land,clear-cut site and farmland in northeast China[J]. Sci Silv Sin, 2013, 49(9):79-88.
[62]丁波,丁贵杰,李先周,等.短期间伐对杉木人工林生态系碳储量的影响[J].中南林业科技大学学报, 2016, 36(8):66-71.DING Bo, DING Guijie, LI Xianzhou, et al. Effects of short term thinning on the carbon storage in Cunninghamia lanceolata plantation ecosystem[J]. J Central South Univer For Technol, 2016, 36(8):66-71.
[63] SETTINERI G, MALLAMACI C, MITROVI'C M, et al. Effects of different thinning intensities on soil carbon storage in Pinus laricio forest of Apennine South Italy[J]. Eur J For Res, 2018, 137(2):131-141.
[64]凌宁.间伐强度对杉木人工林土壤呼吸的影响[D].南京:南京林业大学, 2012.LING Ning. The Impace of Different Thinning Tensity on Soil Respiration in the Chinese fir Plantation[D]. Nanjing:Nanjing Forestry University, 2012.
[65] PAN Ping, ZHAO Fang, NING Jinkui, et al. Impact of understory vegetation on soil carbon and nitrogen dynamic in aerially seeded Pinus massoniana plantations[J]. PLoS One, 2018 1 3(1):e0191952. doi:10.1371/journal.pone.0191952.
[66] WU Jianping, LIU Zhanfeng, CHEN Dima, et al. Understory plants can make substantial contributions to soil respiration:evidence from two subtropical plantations[J]. Soil Biol Biochem, 2011, 43(11):2355-2357.
[67] WANG Xiaoling, ZHAO Jie, WU Jianping, et al. Impacts of understory species removal and/or addition on soil respiration in a mixed forest plantation with native species in southern China[J]. For Ecol Manage, 2011, 261(6):1053-1060.
[68] KUME A, SATOMURA T, TSUBOI N. Effects of understory vegetation on the ecophysiological characteristics of an overstory pine, Pinus densiflora[J]. For Ecol Manage, 2003, 176(1):195-203.
[69] LI Yongfu, JIANG Peikun, CHANG S X, et al. Organic mulch and fertilization affect soil carbon pools and forms under intensively managed bamboo(Phyllostachys praecox)forests in southeast China[J]. J Soils Sediment, 2010, 10(4):739-747.
[70] ZHANG Jiaojiao, LI Yongfu, CHANG S X, et al. Understory vegetation management affected greenhouse gas emissions and labile organic carbon pools in an intensively managed Chinese chestnut plantation[J]. Plant Soil, 2014, 376(1/2):363-375.
[71]吴亚丛,李正才,程彩芳,等.林下植被抚育对樟树人工林土壤活性有机碳库的影响[J].应用生态学报, 2013,24(12):3341-3346.WU Yachong, LI Zhengcai, CHENG Caifang, et al. Effects of understory removal on soil labile organic carbon pool in a Cinnamomum camphora plantation[J]. Chin J Appl Ecol, 2013, 24(12):3341-3346.
[72]李海防,张杏锋.剔除灌草和添加翅荚决明对厚荚相思林土壤温室气体排放的影响[J].应用生态学报, 2010,21(3):563-568.LI Haifang, ZHANG Xingfeng. Soil greenhouse gases emission from an Acacia crassicarpa plantation under effects of understory removal and Cassia alata addition[J]. Chin J Appl Ecol, 2010, 21(3):563-568.
[73] WANG Faming, ZOU Bi, LI Haifang. The effect of understory removal on microclimate and soil properties in two subtropical lumber plantations[J]. J For Res, 2014, 19(1):238-243.
[74] ZHANG Jiaojiao, LI Yongfu, CHANG S X, et al. Understory management and fertilization affected soil greenhouse gas emissions and labile organic carbon pools in a Chinese chestnut plantation[J]. For Ecol Manage, 2015, 337:126-134.
[75] CHATTERJEE S, BANDYOPADHYAY K K, PRADHAN S, et al. Effects of irrigation, crop residue mulch and nitrogen management in maize(Zea mays L.)on soil carbon pools in a sandy loam soil of indo-gangetic plain region[J].Catena, 2018, 165:207-216.
[76] HUANG Zhiqun, XU Zhihong, CHEN Chengrong, et al. Changes in soil carbon during the establishment of a hardwood plantation in subtropical Australia[J]. For Ecol Manage, 2008, 254(1):46-55.
[77]商素云,李永夫,姜培坤,等.天然灌木林改造成板栗林对土壤碳库和氮库的影响[J].应用生态学报, 2012, 23(3):659-665.SHANG Suyun, LI Yongfu, JIANG Peikun, et al. Effects of the conversion from native shrub forest to Chinese chestnut plantation on soil carbon and nitrogen pools[J]. Chin J Appl Ecol, 2012, 23(3):659-665.
[78] MATHERS N J, XU Zhihong. Solid-state13C NMR spectroscopy:characterization of soil organic matter under two contrasting residue management regimes in a 2-year-old pine plantation of subtropical Australia[J]. Geoderma,2003, 114(1/2):19-31.
[79] HUANG Zhiqun, XU Zhihong, CHEN Chengrong. Effect of mulching on labile soil organic matter pools, microbial community functional diversity and nitrogen transformations in two hardwood plantations of subtropical Australia[J].Appl Soil Ecol, 2008, 40(2):229-239.
[80]赵睿宇,李正才,王斌,等.毛竹林地表覆盖年限对土壤有机碳的影响[J].植物生态学报, 2017, 41(4):418-429.ZHAO Ruiyu, LI Zhengcai, WANG Bin, et al. Duration of mulching caused variable pools of labile organic carbon in a Phyllostachys edulis plantation[J]. Chin J Plant Ecol, 2017, 41(4):418-429.
[81]王玉娟.油茶林地覆盖和间种对土壤理化性质及幼林生长的影响[D].长沙:中南林业科技大学, 2009.WANG Yujian. The Effect of Covering and Interplanting on the Soil Physical, Chemical Properties and Growth Condition of Camellia oleifera Young Forest[D]. Changsha:Central South University of Forestry and Technology, 2009.
[82] YOUKHANA A, IDOL T. Tree pruning mulch increases soil C and N in a shaded coffee agroecosystem in Hawaii[J]. Soil Biol Biochem, 2009, 41(12):2527-2534.
[83]郭子武,王为宇,杨清平,等.林地覆盖对雷竹林土壤碳氮磷化学计量特征的影响[J].广西植物, 2013, 33(5):627-632.GUO Ziwu, WANG Weiyu, YANG Qingping, et al. Effect of mulching management on stoichiometry of soil C, N and P in Phyllostachys praecox plantations[J]. Guihaia, 2013, 33(5):627-632.
[84]孙萌,刘洋,李寒,等.有机物覆盖对核桃园土壤有机碳库及酶活性的影响[J].植物营养与肥料学报, 2018, 24(1):270-278.SUN Meng, LIU Yang, LI Han, et al. Effects of organic mulching on soil organic carbon pool and soil enzyme activity in walnut orchard[J]. J Plant Nutr Fert, 2018, 24(1):270-278.